1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola uloloma Brenan

        This species is accepted, and its native range is SE. Kenya to NE. Tanzania.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Type: Tanzania, Pangani District: Bushiri Estate, Faulkner 654 (K!, holo.)
    Habit
    Evergreen small tree, rarely a shrub, (1.5–)3–10(–25) m tall, with a clean trunk and dense, rounded crown; bark pale grey, rough or smooth with longitudinal striations; ultimate branchlets 1 mm thick, dark brown, with numerous smooth raised lenticels of the same colour; bud-scales caducous
    Leaves
    Leaf-blade narrowly elliptic to oblanceolate, sometimes slightly asymmetrical, 4–12 cm long, 1.7–4.7 cm wide, acumen 0.5–1.8 cm long, ± 0.3 cm wide, leathery, 5–7 pairs veins, glabrous above and beneath; petiole terete, 2–5 mm long, 1 mm wide, glabrescent; stipules caducous
    Inflorescences
    Inflorescence in leaf-axils, with fascicles of 1–5 single flowers; bracts 3–4, ± orbicular, the lowest largest, 1.7–2.5 mm long, 2–3 mm across, deeply bilobed, drying with longitudinal ridges, glabrous; pedicel 5–11 mm long, articulated ± midway, puberulous
    Flowers
    Flowers whitish, yellowish or pinkish brown, aromatic, 6–9.5 mm across, divided to within 1.5–2 mm of the base, lobes narrowly triangular, (2.5–)3–4 mm long, 1.2–1.5 mm wide, the upper margins reflexed, white, outer surface sparsely brown stellate-hairy, the hairs 0.1–0.2 mm diameter, inner surface minutely papillate Male flowers with androphore 1.5–2 mm long, 0.2 mm wide, glabrous, anthers 5, uniseriate, forming a ring 1.5–1.8 mm long, 0.4–1.3 mm wide, glabrous Female flowers as the male, but androphore absent, anthers forming a ring at the base of the ovary; ovary of 3–5 carpels, ± spherical, 1.2–2 mm long, 1.8–2.5 mm wide, stigmas ± 0.5 mm long, reflexed
    Male
    Male flowers with androphore 1.5–2 mm long, 0.2 mm wide, glabrous, anthers 5, uniseriate, forming a ring 1.5–1.8 mm long, 0.4–1.3 mm wide, glabrous
    Female
    Female flowers as the male, but androphore absent, anthers forming a ring at the base of the ovary; ovary of 3–5 carpels, ± spherical, 1.2–2 mm long, 1.8–2.5 mm wide, stigmas ± 0.5 mm long, reflexed
    Fruits
    Fruits with a single carpel developed, obliquely obovoid, ± 25 mm long, 17 mm wide, stipe indistinct, rostrum absent, orange, subglabrous
    Ecology
    Coastal forest; 30–500 m
    Conservation
    Of the 13 specimens seen, only five fertile gatherings, listed above, have been seen for Cola uloloma: this species seems rare, poorly known and restricted to a few small coastal forest fragments, mostly the Kaya forests of Kenya. Its extent of occurrence is estimated as less than 20,000 km2. Given that forest quality is declining in several of these patches, and that less than 10 locations are known, Cola uloloma is here assessed as VU B2a,b(iii), i.e vulnerable to extinction.
    Note
    The uniformly short petiole (less than 7 mm) seen in Cola uloloma and C. chlorantha (q.v. for diagnostic characteristics) is unique in East African members of the genus and make these two species easily recognized in the sterile state, assuming that they are recognized as Cola in the first place. Most other species of Cola have a mixture of very long-stalked (to 19 cm) and very short-stalked leaves.   Hawthorne 1645 from the Pugu Hills ( T 6), though sterile, probably represents this species, but without flowers or altitudinal data it is difficult to be certain. Luke (pers. comm.) reported sight records in Kenya for the lower (Kaya Muhaka) and upper (Mangea Hill) altitudinal ranges cited above. Luke (pers. comm.) also reported seeing what was possibly this species, but might also be C. chlorantha (especially the more southern and higher altitudinal specimens) in T 6 (Kimboza, Luke 8814 EA n.v.) and also in T7 ( Luke 8747 EA n.v., 900m alt. and Luke 11311 EA n.v., 1330m alt.).
    Distribution
    Flora districts: K7 T3 T ?6 Range: Not known elsewhere

    Images

    Distribution

    Native to:

    Kenya, Tanzania

    Cola uloloma Brenan appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Faulkner, H.G. [654], Tanzania K000241033 holotype
    Faulkner [654], Tanzania K000241032 holotype

    First published in Kew Bull. 11: 150 (1956)

    Accepted by

    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.

    Literature

    Kew Backbone Distributions
    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    Flora of Tropical East Africa
    • K.T.S.L.: 163 (1994).
    • K.B. 11: 150 (1956)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0