1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola minor Brenan

        This species is accepted, and its native range is SE. Kenya to NE. Tanzania.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Type: Kenya, Kwale District: Marenji Forest, Dale in F.H. 3786 (K!, holo.; EA, iso.)
    Habit
    Evergreen shrub or tree 3–12 m tall, usually single-stemmed.
    Bark
    Bark and wood unknown, sap colourless; ultimate branchlets terete, 0.7–1.5 mm wide, greyish brown, pubescent, becoming puberulous; bud-scales caducous, triangular, ± 2 mm long, puberulous.
    Leaves
    Leaf-blade oblanceolate, rarely narrowly elliptic, 1.9–10(–12) cm long, 0.7–3.6(–4.5) cm wide, apex bluntly acuminate, the acumen ± 1 cm long, base cuneate to acute with convex sides, (7–)8–9 pairs of main veins, leathery, glabrous, dark green drying grey green.
    Petiole
    Petiole terete, 1.5–17(–32) mm long, ± 1 mm wide, puberulous to scurfily brown or grey pubescent when young.
    Inflorescences
    Inflorescences axillary, usually amongst the leaves but also on older wood below, fasciculate with numerous single flowers in bud, but only 1–3 flowers open at one time; bracts 2–3, rounded to oblong, hooded, often bilobed at apex, 1–2.5 mm long, to 1 mm wide, pubescent; stalk 3–4.5(–8) mm long, 0.3–0.4(–0.8) mm wide, with stiff reddish stellate hairs; articulated ± a third of its length from the base
    Flowers
    Flowers white, greenish or yellow, fragrant, perianth divided for 4/5 into (4–)5 partly reflexed lobes, each 2.2–3(–4) mm long, 1.7–2.5(–3) mm wide, outer surface as pedicel, inner surface with finer hairs on upper half of lobes and minutely papillate Male flowers with androphore 1.2–2.2 mm long, 0.1–0.2 mm wide, glabrous; anthers 8–10, uniseriate, glabrous, in a disc 0.5–0.7 mm long, ± 1 mm diameter; ovary vestigial, concealed Female flowers sometimes rather larger than the male, with androphore absent; anthers barely reduced at the base of the ovary; ovary subglobose, slightly trilobed, 1.5–2 mm long, 2.5–3 mm wide, densely tomentose; style ± 1.5 mm long, stigmas 3, recurved
    Male
    Male flowers with androphore 1.2–2.2 mm long, 0.1–0.2 mm wide, glabrous; anthers 8–10, uniseriate, glabrous, in a disc 0.5–0.7 mm long, ± 1 mm diameter; ovary vestigial, concealed
    Female
    Female flowers sometimes rather larger than the male, with androphore absent; anthers barely reduced at the base of the ovary; ovary subglobose, slightly trilobed, 1.5–2 mm long, 2.5–3 mm wide, densely tomentose; style ± 1.5 mm long, stigmas 3, recurved
    Fruits
    Fruits with 1(–3) fruitlets developing, fruitlets obliquely clavate, 8–17 mm long, 10–13 mm wide, apiculus absent, stipe ± 1.5 mm long, orange-yellow, pubescent, with accrescent calyx
    Ecology
    Coastal evergreen forest or thicket; 50–250 m
    Conservation
    Of the 12 specimens known to me of this species, only one is from Tanzania, the remainder being from K 7. It is limited to the severely fragmented remnants of coastal forest, now threatened by clearance. I estimate the area of occurrence of this species to be less than 2000 km2. Accordingly, C. minor is here assessed as VU B2a, b(iii), that is vulnerable.
    Note
    Sometimes confused with Cola pseudoclavata which shares the same habitat and, occasionally, location. C. minor is distinguishable from C. pseudoclavata by its constantly shorter petioles (up to 17(–32) mm long) and dark green uniformly smaller (to 10(–12) cm long) leaf-blades with 8–9 pairs of nerves. All specimens of C. pseudoclavata have petioles up to 50(–60) mm long and yellowish-green leaf-blades with a greater range in size (to 17 cm long) and 5–7(–8) pairs of nerves. C. minor is only known from 10 fertile collections. On present evidence it appears dioecious.
    Distribution
    Flora districts: K7 T3 Range: Not known elsewhere

    Images

    Distribution

    Native to:

    Kenya, Tanzania

    Cola minor Brenan appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1956 Dale, I. R. [3550], Kenya K000241035 Unknown type material
    Jan 1, 1956 Dale, I. R. [3786], Kenya K000241036 holotype
    Verdcourt, B. [1861], Kenya 15375.000

    First published in Kew Bull. 11: 149 (1956)

    Accepted by

    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.

    Literature

    Kew Backbone Distributions
    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    Flora of Tropical East Africa
    • K.T.S.L.: 162, map (1994).
    • K.B. 11: 149 (1956)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0