1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola gigantea A.Chev.

        This species is accepted, and its native range is Benin to Uganda.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Types: Central African Republic, Bondjos, Chevalier 5152 (P, syn., K photo.!); Forêt de Possel, Chevalier 11184 (P, syn.); Dar Banda Oriental, Mbélé, Gounda, Chevalier 7300 (P, syn.; K!, isosyn.)
    Habit
    Deciduous tree 10–35 m tall, with spreading crown.
    Bole
    Bole usually long and straight, sometimes with small, short buttresses.
    Bark
    Bark thick, rough, grey or brown, with deep or slight vertical fissures; slash fibrous, pink to red or mottled yellow, reticulated, turning darker; no exudate recorded.
    Branches
    Flora districts: Ultimate branchlets 5–8 mm thick, densely rusty tomentose with stellate hairs 0.2–0.5 mm across; bud-scales soon falling, triangular, ± 8 mm long, 8 mm wide, rusty tomentose
    Leaves
    Leaf-blade ovate, 15–37(–60) cm long, 10–32(–45) cm wide, apex rounded, base cordate, the internal angle obtuse, ± 120°, rarely truncate, leathery, digitately 5–11-nerved, drying brown or greenish brown above, mature leaves glossy, glabrous, white and densely tomentose beneath; leaf-blades of immature plants larger, ± 45 cm long, 45 cm wide, 3–5-lobed, divided for up to 1/4, subglabrous.
    Petiole
    Petiole terete, 3.5–22 cm long, 0.2–0.35 cm thick, slightly swollen at base and apex, rusty tomentose when young; stipules caducous
    Inflorescences
    Inflorescences borne with the leaves (often on short spur shoots that only produce leaves after flowering), 2–4 per stem, 3–12 cm long, 2–5 cm wide, densely rusty tomentose with stellate hairs; bracts triangular, 1.5–2 mm long; ± 5 branches from the main axis, lowest branch 0.4–3.5 cm long, with 2–4, 1–3flowered partial peduncle; pedicels ± 1.5 mm
    Flowers
    Flowers with perianth white becoming pink, sweetly scented, campanulate, 7–8 mm long, 5.5–10 mm across divided for 1/3 or more into 5 acute, reflexed lobes 2.5–3.5 mm long, 2 mm wide, the margin conspicuously involute; outer surface densely reddish tomentose with minute 8–13-armed stellate hairs, inner surface glabrous in the lower 2.5–3 mm, the upper parts thickly white tomentose with 5–7-armed sinuous stellate hairs Male flowers with androphore 0.7–0.8 mm long, 0.7 mm wide at the base, tapering strongly to the apex, glabrous, but surrounded at the base by long stellate hairs, anther thecae in 2 distinct whorls of 5–7, forming a cylinder 1.5 mm high, 1.5 mm wide, the upper thecae with stellate hairs; vestigial ovary 0.3 mm long, glabrous Female flowers unknown
    Male
    Male flowers with androphore 0.7–0.8 mm long, 0.7 mm wide at the base, tapering strongly to the apex, glabrous, but surrounded at the base by long stellate hairs, anther thecae in 2 distinct whorls of 5–7, forming a cylinder 1.5 mm high, 1.5 mm wide, the upper thecae with stellate hairs; vestigial ovary 0.3 mm long, glabrous
    Female
    Female flowers unknown
    Fruits
    Fruit with follicles obliquely ovoid-ellipsoid, ± 20 cm long, 17 cm high, 6 cm wide, rostrum short and stout, ± 0.7 cm long, stipe absent, thickly covered in minute rusty stellate hairs.
    Seeds
    Seeds oblong-ellipsoid ± 3.5 cm long, 2 cm wide, sessile
    Ecology
    Gallery forest and secondary semi-deciduous forest; 950–1500 m
    Conservation
    Although known from only six specimens in Uganda, this species has such a large range that it is considered of “least concern” for conservation.
    Distribution
    Flora districts: U1 U2 U4 Range: Ghana, Togo, Benin, Nigeria, Cameroon, Central African Republic, CongoKinshasa, Sudan
    [FTEA]
    Use
    Hamilton ( loc. cit.) records that Chimpanzees eat the fruit walls and the ‘arils’.

    Distribution

    Native to:

    Benin, Cameroon, Central African Repu, Chad, Nigeria, Sudan, Uganda, Zaïre

    Cola gigantea A.Chev. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Dec 1, 1953 Maitland, T.D. [1590], Cameroon K000094929
    Dec 1, 1953 Maitland, T.D. [1590], Cameroon K000094930
    Dec 1, 1953 Maitland, T.D. [1590], Cameroon K000094931
    Dec 1, 1953 Maitland, T.D. [1590], Cameroon K000094932

    First published in Mém. Soc. Bot. France 8: 32 (1908)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Lejoly, J. & Geerinck, D. (2010). Flore de la Tshopo (RD Congo): Familles des Malvaceae, Orchidaceae, Sterculiaceae, Tiliaceae Taxonomania 29: 1-41.
    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 1: 1-114. ORSTROM, Bangui.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.

    Literature

    Kew Backbone Distributions
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    Flora of Tropical East Africa
    • Hamilton, Uganda For. Trees: 117 (1981).
    • F.C.B. 10: 315 (1963)
    • K.B. 11: 151 (1956)
    • Bull. Soc. Bot. France: 55: 32 (1908)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0