1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola congolana De Wild. & T.Durand

        This species is accepted, and its native range is DR Congo to SW. Uganda.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Type: Congo-Kinshasa, between Matende and Kibala, Dewèvre s.n. (BR, holo.)
    Habit
    Evergreen, laxly branched tree 3–10 m high.
    Bark
    Bark silvery, brownish grey, flaking in thin papery scales, inner bark pale cream, fibrous, sapwood cream, heartwood darker.
    Stem
    Flora districts: Ultimate stems 3–4 mm thick, silvery grey, with numerous fine ridges, tomentose when very young, often bearing at length epiphytic liverworts and mosses; bud-scales early caducous, narrowly ovate, ± 8.5 mm long, 3 mm wide
    Leaves
    Leaf-blade narrowly elliptic to oblanceolate 7.5–24 cm long, 3–9.5 cm wide, the apex with acumen ± 1.5 cm long, 0.3 cm wide, base cuneate, veins prominent above and below, sparsely tomentose below when young, becoming glabrous on both sides; petiole finely ridged, 0.7–6.5 cm long, 0.1–0.2 cm wide, thinly tomentose when young; stipules caducous
    Inflorescences
    Inflorescences borne below the leaves, with fascicles of 1–20 single flowers per axil, few open at one time, stalks 5–8 mm long, articulated ± 1/3 the way from the base, subglabrous; bracts 4–7, increasing in size towards the apex, wider than long, 1.5–7.5 mm long, 2.5–9.5 mm wide, bilobed for up to half their length, the lowest bracts with rounded lobes, the uppermost acute, becoming perianth-like
    Flowers
    Flowers deep cream, suffused with pink, sometimes yellow or orange, perianth campanulate, 10–14 mm long, 12–18 mm across, divided into (4–)5(–6) triangular lobes each 7–9 mm long, 4–5 mm wide, outside with appressed reddish brown stellate hairs, increasing in density towards the base, and spreading to the apical 1–2 mm of the inner surface, inner surface otherwise glabrous, glossy, with numerous papillae Male flowers with androphore ± cylindrical, 3 mm long, 1 mm wide, glabrous; anthers 8–10, uniseriate, forming a cylinder 1.5 mm long, 2.5–3 mm wide, vestigial carpels 0.6 mm long, glabrous Female flowers usually slightly larger, perianth often 6-lobed, ovary sessile, with a ring of stamens at the base, widely ovoid, 3–4 mm long, 3–4 mm wide, densely villose; style stout, ± 1 mm long; stigmas drying black, elongate, 1–1.5 mm long
    Male
    Male flowers with androphore ± cylindrical, 3 mm long, 1 mm wide, glabrous; anthers 8–10, uniseriate, forming a cylinder 1.5 mm long, 2.5–3 mm wide, vestigial carpels 0.6 mm long, glabrous
    Female
    Female flowers usually slightly larger, perianth often 6-lobed, ovary sessile, with a ring of stamens at the base, widely ovoid, 3–4 mm long, 3–4 mm wide, densely villose; style stout, ± 1 mm long; stigmas drying black, elongate, 1–1.5 mm long
    Fruits
    Fruits with follicles irregularly long ellipsoid to slightly obovoid, 4–10 cm long, 2 cm wide, rostrum 0.5–1 cm long, 0.5 mm wide, stipe absent, glabrous, crimson.
    Seeds
    Seeds spherical to ellipsoid, 1–1.3 cm long
    Ecology
    Evergreen forest; ± 1500 m
    Conservation
    Although known in East Africa only from four specimens, C. congolana is relatively widespread and abundant in the forests of Congo-Kinshasa and so is considered “least concern” for conservation. IUCN (Red List 2000, www.redlist.org) list this species under its synonym C. bracteata De Wild., as VU B1+2c on the fallacious basis that it is endemic to Uganda.
    Distribution
    Flora districts: U2 Range: Congo-Kinshasa

    Distribution

    Native to:

    Uganda, Zaïre

    Cola congolana De Wild. & T.Durand appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Hoyle, A.C. [1403A], Uganda Cola bracteata 15235.000
    Hoyle, A.C. [1403], Uganda Cola bracteata 8868.000

    First published in Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 38: 181 (1899)

    Accepted by

    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Lejoly, J. & Geerinck, D. (2010). Flore de la Tshopo (RD Congo): Familles des Malvaceae, Orchidaceae, Sterculiaceae, Tiliaceae Taxonomania 29: 1-41.
    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.

    Literature

    Flora of Tropical East Africa
    • F.C.B. 10: 292 (1963).
    • B.S.B.B. 38(2): 181 (1899)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0