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  1. Family: Plantaginaceae Juss.
    1. Genus: Veronica L.
      1. Veronica anagallis-aquatica L.

        This species is accepted, and its native range is Temp. Eurasia to Tropical Mountains.

    [FTEA]

    Scrophulariaceae, S.A. Ghazanfar, F.N. Hepper & D. Philcox. Flora of Tropical East Africa. 2008

    Type
    Lectotype: Europe, Herb. Linn. No.5.7 (S) designated by Fischer in F.R. 108: 115 (1977)
    Habit
    Perennial herb usually 30–120 cm high, ± aquatic.
    Stem
    Stems erect, ± quadangular, simple or well-branched, sometimes rooting at the base, hollow, glabrous to sparsely glandular-pubescent
    Leaves
    Leaves linear-lanceolate to ovate-lanceolate, the lower broader than upper ones, ± sessile and amplexicaul, 2–8(–10) cm long, up to 3 cm wide, margins irregularly serrate near the apex, glabrous
    Inflorescences
    Inflorescence lateral with flowers numerous in long racemes at most axils, up to 3 times as long as subtending leaves; pedicels in fruit 3–4 mm long, filiform, spreading or curved upwards, sparsely glandular-pubescent
    Calyx
    Calyx-segments oblong, 3 mm long in fruit, subacute
    Corolla
    Corolla blue with violet veins to white with reddish veins, ± 5 mm in diameter
    Fruits
    Capsule about as long as calyx, elliptical, glabrous, sinus obscure; persistent style much longer than sinus.
    Ecology
    In permanent wet places with shallow running water; 480–2400 m
    Conservation
    Least Concern (LC); widespread and common
    Note
    This species belongs to the Section Beccabunga which consists of a group of similar species, but in East Africa apparently only represented by V. anagallis-aquatica. The species is considerably variable in its vegetative characters probably depending on the wetness of its habitat. This is shown by the gradation from tall, lush plants in rich muddy places to much smaller ones growing in drying mud nearby.
    Distribution
    Range: Widespread in upland tropical Africa, SW Arabia, and almost throughout the northern temperate zones Flora districts: U3 K1 K3 K4 K6 K7 T1 T2 T5
    [FTEA]
    Use
    None recorded on specimens from our area

    Images

    Distribution

    Native to:

    Albania, Algeria, Altay, Assam, Austria, Azores, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bulgaria, Buryatiya, Canary Is., Cape Provinces, Cape Verde, Central European Rus, Chad, China North-Central, China South-Central, China Southeast, Chita, Corse, Cyprus, Czechoslovakia, Denmark, Djibouti, East Aegean Is., East European Russia, East Himalaya, Egypt, Eritrea, Ethiopia, Finland, France, Free State, Germany, Great Britain, Greece, Hungary, Iceland, India, Inner Mongolia, Ireland, Irkutsk, Italy, Kazakhstan, Kenya, Kirgizstan, Krasnoyarsk, Kriti, Krym, KwaZulu-Natal, Lebanon-Syria, Lesotho, Libya, Madeira, Manchuria, Mongolia, Morocco, Myanmar, Namibia, Nepal, Netherlands, North European Russi, Northern Provinces, Northwest European R, Norway, Pakistan, Palestine, Poland, Portugal, Qinghai, Romania, Rwanda, Sardegna, Saudi Arabia, Sicilia, Somalia, South European Russi, Spain, Sudan, Swaziland, Sweden, Switzerland, Tadzhikistan, Tanzania, Tibet, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Tuva, Uganda, Ukraine, Uzbekistan, West Himalaya, West Siberia, Xinjiang, Yemen, Yugoslavia, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Alabama, Alberta, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Bolivia, Brazil South, Brazil Southeast, British Columbia, California, Chile Central, Chile North, Chile South, Colorado, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Juan Fernández Is., Kansas, Kentucky, Korea, Maine, Mauritius, Michigan, Missouri, Montana, Nebraska, Nevada, New Jersey, New Mexico, New York, New Zealand North, North Carolina, North Dakota, Ohio, Ontario, Oregon, Pennsylvania, Peru, Primorye, Rhode I., South Dakota, St.Helena, Tennessee, Texas, Uruguay, Utah, Vermont, Virginia, Washington, West Virginia, Wisconsin, Wyoming

    Accepted Infraspecifics

    Synonyms

    Other Data

    Veronica anagallis-aquatica L. appears in other Kew resources:

    Bibliography

    First published in Sp. Pl.: 12 (1753)

    Accepted by

    • Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
    • Arbo, M.M. & al. (2018). Flora Argentina. Flora vascular de la República Argentina 20(1): 1-488. INTA, IMBIV & IBODA.
    • Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
    • Sennikov, A.N. (ed.) (2017). Flora of Uzbekistan 2: 1-200. Navro'z Publishers, Toshkent.
    • Werier, D. (2017). Catalogue of the Vascular plants of New York state Memoirs of the Torrey Botanical Club 27: 1-542.
    • Kliment, J., Turis, P. & Janišová, M. (2016). Taxa of vascular plants endemic to the Carpathian Mts Preslia. Casopsi Ceské Botanické Spolecnosti 88: 19-76.
    • Mill, R. & al. (2015). Flora of Pakistan 220: 1-331. Department of Botany, University of Karachi, Karachi.
    • Gilman, A.V. (2015). New flora of Vermont Memoirs of the New York Botanical Garden 110: 1-614.
    • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
    • Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
    • Larson, J., Reif, B., Nelson, B.E. & Hartman, R.L. (2014). Floristic studies in North Central New Mexico, U.S.A. the Sange de Cristo mountains Journal of the Botanical Research Institute of Texas 8: 271-303.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
    • Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
    • Kumar, S. (2012). Herbaceous flora of Jaunsar-Bawar (Uttarkhand), India: enumerations Phytotaxonomy 12: 33-56.
    • Llewellyn, O.A., Hall, M., Miller, A.G., Al-Abbasi, T.M., Al-Wetaid, A.H., Al-Harbi, R.J. & Al-Shammari, K.F. (2011). Important plant areas in the Arabian peninsula: 4. Jabal Aja Edinburgh Journal of Botany 68: 199-224.
    • Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
    • Authier, P. & Covillot, J. (2011). Catalogue actualisé des plantes de l'île de Rhodes (Grèce) Saussurea; Travaux de la Société Botanique de Genève 41: 131-170.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Musselman, L.J. (2011). Checklist of Plants of Lebanon and Syria http://ww2.odu.edu/~lmusselm/plant/lebsyria/Checklist%20of%20Lebanon%20Plants.pdf.
    • Ghazanfar, S.A., Hepper, F.N. & Philcox, D. (2008). Flora of Tropical East Africa, Scrophulariaceae: 1-211.
    • Rodríguez, R., Grau, J., Baeza, C. & Davies, A. (2008). Lista comentada de las plantas vasculares de los Nevados de Chillan, Chile Gayana. Botánica 65: 153-197.
    • Singh, M. & Ali, S.J. (2008). A study on aquatic Angiosperms of district Mau (East U.P.) Journal of East African Natural History 32: 877-890.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Kamilya, P. (2008). Survey of weed flora of Atrai river bed in Dakshin-Dinajpur in District of West Bengal, India Pleione 2: 77-86.
    • Polozhij, A.V. & Peschkova, G.A. (eds.) (2007). Flora of Siberia 12: 1-221. Scientific Publishers, Inc., Enfield, Plymouth.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Aver'ianov, L.V. & al. (2006). Illyustrirovannyi opredelitel' rastenii Leningradskoi oblasti: 1-799. Nauka, Moskva.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Jones, R.L. (2005). Plant life of Kentucky. An illustrated guide to the vascular flora: 1-833. The universitry press of Kentucky.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
    • Grierson, A.J.C. & Long, D.G. (2001). Flora of Bhutan 2: 1-1675. Royal Botanic Gardens, Edinburgh.
    • Bosser, J. & al. (eds.) (2000). Flore des Mascareignes 127-135: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Fischer, E. (1999). Flore d'Afrique Centrale (Zaire-rwanda-Burundi), Scrophulariaceae: 1-217. Jardin Botanique National de Belgique.
    • Z.Wu & P.H.Raven (eds.) (1998). Flora of China 18: 1-449. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. Missouri Botanical Garden.
    • Barkalov, V.Y. (ed.) (1991). Plantae Vasculares Orientalis Extremi Sovietici 5: 1-388. Nauka, Leningrad.
    • Meikle, R.D. (1985). Flora of Cyprus 2: 833-1970. The Bentham-Moxon Trust Royal Botanic Gardens, Kew.
    • Troupin, G. (ed.) (1985). Flora du Rwanda 3: 1-729. Musee Royal de l'Afrique Centrale.
    • Chukavina, A.P. (ed.) (1984). Flora Tadzhikskoi SSR 7: 1-562. Izd-vo Akademii nauk SSSR, Moskva.
    • Davis, P.H. (ed.) (1978). Flora of Turkey and the East Aegean Islands 6: 1-825. Edinburgh University Press, Edinburgh.
    • Tutin, T.G. & al. (eds.) (1972). Flora Europaea 3: 1-370. Cambridge University Press.
    • Vvedensky, A.I. (ed.) (1961). Flora Uzbekistana 5: 1-667. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.

    Literature

    Kew Backbone Distributions

    • Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
    • Flora of North America Editorial Committee (2019). Flora of North America North of Mexico 17: 1-737. Oxford University Press, New York, Oxford.
    • Werier, D. (2017). Catalogue of the Vascular plants of New York state Memoirs of the Torrey Botanical Club 27: 1-542.
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Mill, R. & al. (2015). Flora of Pakistan 220: 1-331. Department of Botany, University of Karachi, Karachi.
    • Gilman, A.V. (2015). New flora of Vermont Memoirs of the New York Botanical Garden 110: 1-614.
    • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
    • Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Larson, J., Reif, B., Nelson, B.E. & Hartman, R.L. (2014). Floristic studies in North Central New Mexico, U.S.A. the Sange de Cristo mountains Journal of the Botanical Research Institute of Texas 8: 271-303.
    • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
    • Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
    • Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Marhold, K. (ed.) (2012). IAPT/IOPB chromosome data 13 Taxon 61: 889-902.
    • Llewellyn, O.A., Hall, M., Miller, A.G., Al-Abbasi, T.M., Al-Wetaid, A.H., Al-Harbi, R.J. & Al-Shammari, K.F. (2011). Important plant areas in the Arabian peninsula: 4. Jabal Aja Edinburgh Journal of Botany 68: 199-224.
    • Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
    • Authier, P. & Covillot, J. (2011). Catalogue actualisé des plantes de l'île de Rhodes (Grèce) Saussurea; Travaux de la Société Botanique de Genève 41: 131-170.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Musselman, L.J. (2011). Checklist of Plants of Lebanon and Syria http://ww2.odu.edu/~lmusselm/plant/lebsyria/Checklist%20of%20Lebanon%20Plants.pdf.
    • Kosachev, P.A. (2010). Synopsis of the families Scrophulariaceae Juss. and Pediculariaceae Juss. of Altai mountain country Turczaninowia 13(1): 19-102.
    • Hadinec, J. & Lustyk, P. (2009). Additions to the flora of the Czech Republic. VIII. Zprávy Ceské Botanické Spolecnosti 44: 185-319.
    • Rodríguez, R., Grau, J., Baeza, C. & Davies, A. (2008). Lista comentada de las plantas vasculares de los Nevados de Chillan, Chile Gayana. Botánica 65: 153-197.
    • Singh, M. & Ali, S.J. (2008). A study on aquatic Angiosperms of district Mau (East U.P.) Journal of East African Natural History 32: 877-890.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Polozhij, A.V. & Peschkova, G.A. (eds.) (2007). Flora of Siberia 12: 1-221. Scientific Publishers, Inc., Enfield, Plymouth.
    • Aver'ianov, L.V. & al. (2006). Illyustrirovannyi opredelitel' rastenii Leningradskoi oblasti: 1-799. Nauka, Moskva.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Jones, R.L. (2005). Plant life of Kentucky. An illustrated guide to the vascular flora: 1-833. The universitry press of Kentucky.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
    • Grierson, A.J.C. & Long, D.G. (2001). Flora of Bhutan 2: 1-1675. Royal Botanic Gardens, Edinburgh.
    • Bosser, J. & al. (eds.) (2000). Flore des Mascareignes 127-135: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Fischer, E. (1999). Flore d'Afrique Centrale (Zaire-rwanda-Burundi), Scrophulariaceae: 1-217. Jardin Botanique National de Belgique.
    • Z.Wu & P.H.Raven (eds.) (1998). Flora of China 18: 1-449. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. Missouri Botanical Garden.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Barkalov, V.Y. (ed.) (1991). Plantae Vasculares Orientalis Extremi Sovietici 5: 1-388. Nauka, Leningrad.
    • Meikle, R.D. (1985). Flora of Cyprus 2: 833-1970. The Bentham-Moxon Trust Royal Botanic Gardens, Kew.
    • Troupin, G. (ed.) (1985). Flora du Rwanda 3: 1-729. Musee Royal de l'Afrique Centrale.
    • Chukavina, A.P. (ed.) (1984). Flora Tadzhikskoi SSR 7: 1-562. Izd-vo Akademii nauk SSSR, Moskva.
    • Brown, L.C. (1982). The Flora and Fauna of St Helena: 1-88. Land Resources Development Centre, Surbiton, England.
    • Davis, P.H. (ed.) (1978). Flora of Turkey and the East Aegean Islands 6: 1-825. Edinburgh University Press, Edinburgh.
    • Tutin, T.G. & al. (eds.) (1972). Flora Europaea 3: 1-370. Cambridge University Press.
    • Vvedensky, A.I. (ed.) (1961). Flora Uzbekistana 5: 1-667. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
    • Schischkin, B.K. & al. (eds.) (1954). Flora Turkmenii 6: 1-402. Turkmenskoe gosudarstvennoe izd., Ashkhabad.

    Flora of Tropical East Africa

    • Fl. Ethiop. & Eritr. 5: 283 (2006).
    • Fischer, F.A.C. Scrophulariaceae: 206 (1999)
    • U.K.W.F.: 258, pl. 111 (1994)
    • F.Z.: 8(2): 80 (1990)
    • Fl. Europ. 3: 248 (1972)
    • F.T.A. 4(2): 357 (1906) as V. anagallis
    • Sp. Pl. :12 (1753)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Science Photographs
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/