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This species is accepted, and its native range is Nepal to S. Central China and Indo-China.
A specimen from Kew's Herbarium

[KBu]

Suddee, S. & Paton, A. 2009. A revision of Anisochilus Wall. ex Benth. (Lamiaceae). Kew Bulletin 64: 235. DOI: https://doi.org/10.1007/s12225-009-9115-5

Conservation
Widespread, Least Concern.
Distribution
Nepal, Eastern India (Sikkim, Assam), Bhutan, Burma, SW China (Yunnan), Thailand, Laos and Vietnam. Nepal, Bhutan, India, Burma, SW China (Yunnan), Thailand, Laos and Vietnam.
Ecology
Edges of forest or road sides in primary evergreen forest, lower montane forest, moist mixed deciduous forest; 300 – 1900 m. Flowering October – January, fruiting December – April. In grass fields, primary evergreen forest, lower montane forest, moist mixed deciduous forest; 900 - 1900 m.
Morphology General Habit
Erect annual herbs up to 1.50 m tall Erect annual herbs up to 1.50 m tall
Morphology Leaves
Leaves petiolate, opposite decussate, often with axillary young leaves, elliptic-oblong or oblong-lanceolate, 30 – 160 × 15 – 70 mm, apex acuminate, base attenuate, margin dentate-serrate, membranous, glabrescent to sparsely pubescent above, glabrescent to pubescent with sessile glands beneath, lateral veins 5 – 12; petiole slender, up to 50 mm long, pubescent, with sparse sessile glands, often with 2 rows of longer hairs along edge near leaf base Leaves opposite, often with axillary young leaves, membranous, elliptic. oblong or oblong-lanceolate, 30 - 150 x 15 - 70 mm, apex acuminate, base attenuate, margin dentate, never revolute, glabrescent to sparsely pubescent above, glabrescent to pubescent with sessile glands beneath, hairs much denser on midrib both sides, lateral veins 5 - 12, reticulate venation prominent beneath; petiole distinct, slender, up to 50 mm long, pubescent with sparse sessile glands, often with 2 rows of longer hairs along edge near leaf base
Morphology Reproductive morphology Flowers Calyx
Calyx ovoid, c. 1 mm long at anthesis, densely villous; fruiting calyx obliquely 5-toothed, 4 – 6 mm long; uppermost tooth subulate, beak-like, deflexed after anthesis and concealing throat, pubescent; lateral 4 oblique teeth minute, erect; tube ventrally saccate around the middle, constricted at throat, villous Calyx ovoid, c. 1 mm long at anthesis, densely villous outside; fruiting calyx obliquely 5-toothed; posterior lip 1-lobed, subulate, beak-like, 1.5 - 3 mm long, deflexed after anthesis and concealing throat, pubescent with or without sessile glands outside; anterior lip with 4 oblique and erect minute teeth, with 2 teeth on each side at the base of posterior calyx lip, seemingly like the lateral teeth of posterior lip, the outer ones very minute, sometimes obscure; tube 4 - 6 mm long, ventrally saccate around the middle, constricted at throat, villous outside
Morphology Reproductive morphology Flowers Corolla
Corolla white to bluish-purple, 4 – 6 mm long, villous, with spare sessile glands Corolla white to bluish-purple, 4 - 6 mm long; posterior lip shortly 4-lobed, erect, the median lobes orbicular, back of lobes sparsely villous with sessile glands, lateral lobes very minute; anterior lip obovate-oblong, concave, constricted at base, much longer than posterior, villous with sessile glands outside, glabrous inside; tube abruptly expanded towards throat from midpoint, base narrowed into a slender cylindrical tube, villous with spare sessile glands outside, glabrous inside
Morphology Reproductive morphology Fruits
Nutlets dark brown to black, oblong or ellipsoid, 1 – 1.2 × 0.6 – 0.8 mm Nutlets dark brown to black, oblong, 0.6 - 0.8 x 1 - 1.2 mm.
Morphology Reproductive morphology Inflorescences
Inflorescence terminal and axillary, simple or branched; spike-like head often narrowed toward apex in bud and becoming cylindric after anthesis, up to 55 mm long and 12 mm wide; bracts narrowly lanceolate, pubescent Inflorescence simple or branched; spikes often narrowed toward apex in bud and becoming cylindric after anthesis, up to 55 mm long and 10 mm wide, flowers dense; peduncle long and slender; bracts narrowly lanceolate, up to 2.5 mm long and 0.5 mm wide, apex acuminate, base sessile, abaxial pubescent with sessile glands, adaxial glabrous with prominent median longitudinal nerve, ciliate, often forming a small apical coma
Morphology Stem
Stems round-quadrangular or quadrangular, usually 4-longitudinal grooved when dry, glabrescent to pubescent, with sessile glands, often with purple spots Stems branched, round-quadrangular or quadrangular, glabrescent to pubescent with short retrose or subpatent hairs, that become longer at nodes and denser on the inflorescence axis, with or without sessile glands, old stem glabrous or glabrescent
Note
Bentham, in his original description, described Anisochilus pallidus as having an obliquely 5-toothed fruiting calyx; he also described the upper subulate calyx lip as the posterior lip and the other 4 oblique minute teeth as the anterior lip. By contrast Hooker in Fl. Brit. India 4: 629 (1885) described the 3 upper teeth as the posterior calyx lip and did not mention the remaining 2 minute teeth. Bentham's original description is in agreement with this account. Most continental SE Asian specimens have an anterior lip with 4 minute teeth but these are arranged obliquely at the base of the posterior one.
Phenology
Flowering November -January, fruiting December - April.
Type
Burma, Taong Dong, 26 Nov. 1826, Wallich Cat. 2754, [lectotype K! (Herb. Benthamianum); isolectotypes G-DC (microfiche!), K-W!]. Burma, Taong Dong, 26 Nov. 1826, Wallich Cat. 2754, [K! (Herb. Benthamianum), lectotype chosen here; isolectotypes G-DC (microfiche!), K-W!].
Vernacular
Indian (Assam): Hneminuchu. Vietnamese: Dithan tai. Vietnamese: Dithan tai.

Native to:

Assam, Bangladesh, China South-Central, East Himalaya, Laos, Myanmar, Nepal, Thailand, Vietnam

Coleus pallidus (Wall. ex Benth.) A.J.Paton appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 1999 Wallich, N. [9754], Myanmar Anisochilus pallidus K000674759 lectotype

First published in PhytoKeys 129: 81 (2019)

Accepted by

  • Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. (2021). The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. https://doi.org/10.1038/s41597-021-00997-6 Scientific Data 8: 215.

Literature

Kew Backbone Distributions

  • Khanam, M. & Hassan, M.A. (2008). Lamiaceae Flora of Bangladesh 58: 1-161. Bangladesh National Herbarium, Dhaka.

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
Kew Bulletin
http://creativecommons.org/licenses/by-nc-sa/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0