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This species is accepted, and its native range is Indo-China.

[KBu]

Suddee, S. & Paton, A. 2009. A revision of Anisochilus Wall. ex Benth. (Lamiaceae). Kew Bulletin 64: 235. DOI: https://doi.org/10.1007/s12225-009-9115-5

Conservation
Least concern (Suddee et al.2004: 384).
Distribution
Cambodia, Laos and Thailand. Cambodia, Laos and Thailand.
Ecology
In open grassy ground, marshy areas, dipterocarp forest, mixed-deciduous forest; 20 - 870 m. On rocks in open grassy places in Dipterocarp forest or mixed-deciduous forest, in marshy areas; 20 – 870 m. Flowering September – February, fruiting October – March.
Morphology General Habit
Erect herbs or undershrub 1- 1.50 m tall Erect short-lived perennial herbs or undershrub 1 – 1.5 m tall
Morphology Leaves
Leaves opposite, often pseudo-verticillate due to the presence of axillary young leaves, lanceolate, 15 - 90 x 5 - 25 mm, apex acute, base attenuate, margin crenate, serrate or subentire, often revolute, scabrate to tomentose with or without sessile glands above, tomentose with sessile glands beneath, sometimes glands hardly visible because of dense indumentum, lateral veins 3 - 6, prominent beneath; petiole absent to 10 mm long, pubescent Leaves petiolate or sessile, opposite decussate, often pseudo-verticillate due to the presence of axillary young leaves, lanceolate, 15 – 90 × 5 – 25 mm, apex acute, base attenuate, margin crenate, serrate or subentire, often revolute, scabrate to tomentose with or without sessile glands above, tomentose with sessile glands beneath, sometimes glands hardly visible because of dense indumentum, lateral veins 3 – 6, prominently raised beneath; petiole 0 – 10 mm long, pubescent
Morphology Reproductive morphology Flowers Calyx
Calyx ovoid, 1 - 1.2 mm long at anthesis, tube densely villous outside; fruiting calyx bilabiate; posterior lip 1-lobed, oblong, 0.8 - 1.2 x 1 - 1.2 mm, slightly recurved, not concealing throat, apex obtuse or rounded, occasionally shallowly emarginate or shortly apiculate, villous to densely villous with sessile glands outside; anterior lip shortly 4-lobed, subequal, apex obtuse or rounded, deflexed or not; tube 3 - 5 mm long, ventrally saccate below the middle, villous outside Calyx ovoid, 1 – 1.2 mm long at anthesis, tube densely villous; fruiting calyx 3 – 6 mm long; uppermost tooth oblong, slightly decurved, not concealing throat, apex obtuse or rounded, occasionally shallowly emarginate or shortly apiculate, villous with sessile glands; 4 remaining teeth subequal, apex obtuse or rounded, reflexed or not, arranged obliquely on throat at base of uppermost tooth; tube ventrally saccate below the middle, villous
Morphology Reproductive morphology Flowers Corolla
Corolla white, whitish-purple or purple, 8 - 10 mm long; posterior lip shortly 4-lobed, subequal, orbicular, back of lobes villous with sparse sessile glands; anterior lip oblong or obovate-oblong, slightly concave, longer than posterior, back of lobe villous at the middle, edge thin and glabrous; tube gradually dilated towards throat from base and dorsally decurved around midpoint, villous with sessile glands outside, glabrous inside Corolla white, whitish-purple or purple, 8 – 10  mm long, villous with sessile glands
Morphology Reproductive morphology Fruits
Nutlets dark brown or black, ovoid, c. 1 mm long. Nutlets dark brown or black, ovoid, c. 1 mm long
Morphology Reproductive morphology Inflorescences
Inflorescence much branched; spikes cylindric, 10 - 150 x 10 - 12 mm, flowers dense; bracts ovate or narrowly so, up to 3 mm long and 2.5 mm wide, acuminate, concave, sessile, abaxially pubescent with sessile glands, adaxially glabrous to pubescent Inflorescence terminal, much branched; spike-like head cylindric, 10 – 150 × 10 – 12 mm; bracts ovate or lanceolate, pubescent, with sessile glands
Morphology Stem
Stems woody or herbaceous and woody at base, quadrangular, hoary-pubescent with short antrose hairs, with or without sparse sessile glands, glands varying from yellow to red, old stem glabrescent Stems woody or herbaceous and woody at base, quadrangular, hoary-pubescent
Note
Anisochilus harmandii Doan (1936: 943, f. 99, 11 – 13), nom. inval.; Murata (1976: 179).
Phenology
Flowering October - February; fruiting October - March.
Type
Cambodia, Mlu Prey, Jan. 1876, Harmand 326 (holotypus P!; isotypi A!, P!). Cambodia, Mlu Prey, Jan. 1876, Harmand 326 (holotype P!; isotypes A!, P!).
Vernacular
Thai: Khuu muai (Chong-Trat), Khruu muai (Trat). Thai: Khuu muai (Chong-Trat), Khruu muai (Trat).

Native to:

Cambodia, Laos, Thailand

Coleus harmandii (Doan ex Suddee & A.J.Paton) A.J.Paton appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Feb 2, 2011 Phuong, V.X. et al. [HNK 330], Vietnam Anisochilus harmandii K000249863

First published in PhytoKeys 129: 56 (2019)

Accepted by

  • Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. (2021). The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. https://doi.org/10.1038/s41597-021-00997-6 Scientific Data 8: 215.

Literature

Kew Backbone Distributions

  • Suddee, S. & Paton, A. (2009). A revision of Anisochilus Wall. ex Benth. (Lamiaceae) Kew Bulletin 64: 235-257.

Herbarium Catalogue Specimens

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
Kew Bulletin
http://creativecommons.org/licenses/by-nc-sa/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0