1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola pseudoclavata Cheek

        This species is accepted, and its native range is SE. Kenya to E. Tanzania.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Type: Tanzania, Tanga District: Kivomila River, 8 km S of Ngomeni, Drummond and Hemsley 3519 (K!, sheet 2, holo.; EA, K! sheet 1, iso.)
    Habit
    Evergreen tree or shrub 5–21 m tall.
    Bole
    Bole sometimes buttressed at the base, bark smooth, pale grey or brown, wood hard, pale yellow or white; ultimate branchlets 1.5–3 mm thick, glabrous, smooth, whitish, becoming shallowly longitudinally ridged; bud-scales caducous.
    Leaves
    Leaf-blade ovate to oblanceolate, 2.5–17 cm long, 1.1–6.1 cm wide, apex acuminate, base cuneate to obtuse, sometimes rounded or slightly cordate, leathery, 5–8-nerved on each side of the midrib, finely reticulate, yellowish or lime green, drying brownish or greyish green, glabrous; petiole ± terete, 0.1–5(–6.5) cm long, ± 1 mm wide, slightly swollen at base and apex, glabrous; stipules caducous
    Inflorescences
    Inflorescence axillary, amongst the leaves and on older wood below, fasciculate with numerous single flowers in bud, 1–3 flowers open at one time; bracts 3–6, ± elliptic, 1–2 mm long, entirely basal or inserted up to halfway along the pedicel, 1–2 mm long, increasing in size towards the flower, cucullate, entire or bilobed, puberulous; pedicel 4–6.5 mm long, 0.2–0.3 mm wide, with short-armed reddish stellate hairs; articulation basal, inconspicuous
    Flowers
    Flowers with perianth cream inside, outside with brown hairs, scent not recorded, divided for 4/5 into 6–8(–10) ± patent lobes, each (3.2–)3.5–4 mm long, 1–1.8 mm wide, the margins reflexed; outer surface as pedicel, inner with a few stellate hairs at tip, and minutely papillate Male flowers with androphore (2.5–)3–4 mm long, 0.1–0.2 mm wide, glabrous, anthers uniseriate, 8–10, glabrous, in a disc ± 1 mm long, 1–1.5 mm diameter, cream or orange; ovary vestigial, concealed. Female flowers with androphore absent; anthers only slightly reduced at base of ovary; ovary depressed globose, 1–1.5 mm long, 1.5–2 mm diameter, strongly 5-lobed, bristly tomentose, cream; style 1–1.5 mm long
    Male
    Male flowers with androphore (2.5–)3–4 mm long, 0.1–0.2 mm wide, glabrous, anthers uniseriate, 8–10, glabrous, in a disc ± 1 mm long, 1–1.5 mm diameter, cream or orange; ovary vestigial, concealed.
    Female
    Female flowers with androphore absent; anthers only slightly reduced at base of ovary; ovary depressed globose, 1–1.5 mm long, 1.5–2 mm diameter, strongly 5-lobed, bristly tomentose, cream; style 1–1.5 mm long
    Fruits
    Fruitlets 1–3, ripening yellow and red, ± obliquely clavate, 1.4–1.9 cm long, 1.2–1.8 cm wide, apiculus oblique, inconspicuous, ± 1 mm long, covered in fine golden stellate hairs
    Ecology
    Riverine or coastal evergreen or semi-deciduous forest; 30–600 m
    Conservation
    Flora districts: Known from ten fertile specimens in Kenya, four in Tanzania and numerous sterile specimens (identification therefore uncertain), C. pseudoclavata is fairly common within a fairly large range and appears not to be of immediate concern for conservation, although threats to its forest habitat could change this. For the moment it is rated as “near threatened”.
    Note
    Fruitlet width: 0.9–1 cm Seed orientation in fruitlet: Longitudinal Fruitlet stipe: Well-defined, ± 10 mm long, width even, 2 mm Perianth lobe number: 5 Indumentum of outer perianth & pedicel: Long-armed white stellate hairs Cola pseudoclavata Fruitlet width: 1.2–1.8 cm Seed orientation in fruitlet:Transverse Fruitlet stipe: Poorly defined, ± 7 mm long, dilating steadily from pedicel to fruit body, ± 4 mm wide at midpoint Perianth lobe number: 6–8(–10) Indumentum of outer perianth & pedicel: Short-armed red stellate hairs Dioecious on the evidence of Drummond & Hemsley 3519 & 3520. The former specimen is the only one known to have female flowers. Cola pseudoclavata is sometimes confused with C. minor in coastal Kenya, which is distinguished vegetatively by its smaller, dark green leaves with more uniformly short leaf-stalks. Sometimes confused in Tanzania with C. discoglypremnophylla (for differences, see there).  Hitherto known in the FTEA area as C. clavata following Brenan (loc. cit. 1958). The type of C. clavata is a fruiting collection from Mozambique. Brenan, in attempting to work out the correct name for this FTEA taxon, (then known only from flowering material), concluded that it was C. clavata on the basis of his observations of perianth fragments persisting at the base of the fruits on the Mozambiquan type. However Wild (in litt.) in working out the Cola species of the Flora Zambesiaca region doubted Brenan’s conclusion and postulated that the FTEA taxon differed and required a new name. In the intervening 40 years ample specimens have now accumulated of what are clearly two morphologically distinct and geographically disjunct species. These are easily distinguished using the characters in the table below. Accordingly a new name has been provided for the FTEA taxon above. Cola clavata
    Distribution
    Flora districts: K7 T3 T6
    [FTEA]
    Use
    Wood used as handles for Jembes and axes, Mahunda 1. Choice firewood, Medley 344.

    Distribution

    Doubtfully present in:

    Somalia

    Native to:

    Kenya, Tanzania

    Cola pseudoclavata Cheek appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Nov 1, 2007 Luke, W.R.Q. [10723], Kenya K000197459

    First published in Fl. Trop. E. Africa, Sterculiac.: 43 (2007)

    Accepted by

    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.

    Literature

    Kew Backbone Distributions
    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0