1. Family: Malvaceae Juss.
    1. Genus: Cola Schott & Endl.
      1. Cola ruawaensis Cheek

        This species is accepted, and its native range is S. Tanzania.

    [FTEA]

    Sterculiaceae, Martin Cheek & Laurence Dorr; Nesogordonia, Laurence Dorr, Lisa Barnett. Flora of Tropical East Africa. 2007

    Type
    Typus: Tanzania, Lindi District: Ruawa forest, Mbago et al. 2279 (K!, holo.; DAR, iso.)
    Habit
    Shrub 5 m tall; leafy stems white, longitudinally ridged, 3 mm wide, lenticels inconspicuous; current year’s growth pale brown, tomentose, glabrous at length; bud-scales caducous, not seen
    Leaves
    Leaf-blade simple, elliptic (smallest leaves) to obovate (larger), 7.5–16 cm long, 3.5–7 cm wide, acumen 0.3–1 cm long, base rounded to cordate (smallest leaves) or acute, lateral nerves 9 on each side of the midrib, prominent; petiole 0.5–6.5 cm long, 1 mm wide, the base and apex distinctly swollen, covered in pale brown tomentum, glabrescent; stipules caducous, not seen. Inflorescence with flowers borne with the leaves, axillary, in fascicles of 2–5, sessile, bracts ± 6 per flower and enveloping it in bud, papery, increasing in size with proximity to flower, innermost bract, suborbicular, ± 5 mm long, 8 mm wide, apex retuse, outer surface yellow-brown tomentose, apex, margins and inner surface glabrous, dark brown
    Flowers
    Male flowers yellow, with brownish hairs, 20–25 mm across, perianth widely campanulate, divided to within 7–8 mm of the base into 6 or 7 lobes, lobes oblong-triangular, 12 mm long, 5 mm wide, apex obtuse, margin truncate, inner surface evenly papillate over the entire surface, outer surface brown tomentose, with two types of hair: short hairs dense, arms erect, long hairs scattered, arms appressed, 5; androphore conical-cylindrical, 5.5 mm long, 3.5 mm wide at base, 1.8 mm wide at apex, indumentum simple, patent, moderately dense; anthers uniseriate, 15; anther-head 2.5 mm long, 6.5 mm wide Female flowers and fruits unknown
    Male
    Male flowers yellow, with brownish hairs, 20–25 mm across, perianth widely campanulate, divided to within 7–8 mm of the base into 6 or 7 lobes, lobes oblong-triangular, 12 mm long, 5 mm wide, apex obtuse, margin truncate, inner surface evenly papillate over the entire surface, outer surface brown tomentose, with two types of hair: short hairs dense, arms erect, long hairs scattered, arms appressed, 5; androphore conical-cylindrical, 5.5 mm long, 3.5 mm wide at base, 1.8 mm wide at apex, indumentum simple, patent, moderately dense; anthers uniseriate, 15; anther-head 2.5 mm long, 6.5 mm wide
    Female
    Female flowers and fruits unknown
    Figures
    Fig. 5, p 35
    Ecology
    Lowland evergreen forest dominated by Scorodophloeus on coral rag; ± 330 m
    Conservation
    Cola ruawaensis is only known from the type locality. Ruawa forest is threatened by agricultural encroachment and illegal logging according to Clarke, P. (1995) Status Reports for 6 Coastal Forests in Lindi Region, Tanzania. Accordingly the taxon is here assessed as CR B2a,b(iii), critically endangered.
    Note
    Cola ruawaensis is closely related to C. octoloboides of Kenya, also restricted to limestone, and only known from 1 or 2 sites. It differs in the non-persistent stipules, the deeply retuse (not rounded) bracts, the perianth divided to 7–8 mm from the base (not 2–5 mm), and the androphore with simple hairs (not stellate hairs). More collections are needed to complete our knowledge of the species.
    Distribution
    Flora districts: T8 Range: Not known elsewhere

    Images

    Distribution

    Native to:

    Tanzania

    Cola ruawaensis Cheek appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 2005 Mbago, F.M. [FMM2279], Tanzania K000691489 holotype

    First published in Fl. Trop. E. Africa, Sterculiac.: 34 (2007)

    Accepted by

    • Cheek, M. & Dorr, L. (2007). Flora of Tropical East Africa, Sterculiaceae: 1-134.

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0