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This genus is accepted, and its native range is Kenya to S. Africa, Madagascar, Indo-China.
Karomia speciosa

[LKGF]

Harley, R.M. et al. (2004). Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Springer-Verlag, Berlin, Heidelberg, New York.

Morphology General Habit
Trees and shrubs, at least some aromatic
Morphology Leaves
Leaves opposite, petiolate, ovate to obovate or suborbicular, entire or crenate
Morphology Reproductive morphology Inflorescences
Inflorescence a lax axillary cyme or terminal thyrse
Morphology Reproductive morphology Flowers Calyx
Calyx actinomorphic, greatly accrescent, broadly infundibular in fruit, 5-lobed, lobes ovate to dome-shaped or obsolescent
Morphology Reproductive morphology Flowers Corolla
Corolla purple or whitish, 1-lipped with only anterior lip present, 5-lobed, anterior lobe largest, concave, sometimes fimbriate, tube straight or curved, sometimes gibbous, corolla in bud usually asymmetrical
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, subequal, long-exserted, filaments arched, anther-thecae parallel or divaricate, confluent or separate at dehiscence
Morphology Reproductive morphology Flowers Pollen
Pollen psilate or bearing blunt, irregularly shaped supratectal protuberances
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary unlobed
Morphology Reproductive morphology Flowers Gynoecium Stigma
Stigma-lobes equal or unequal
Morphology Reproductive morphology Flowers Disc
Disc poorly developed to absent
Morphology Reproductive morphology Fruits
Fruit dry, obconical and apically truncate or with four horn-like projections, or globose, usually pubescent, 4-seeded, remaining intact or splitting into four mericarps, attachment scar as long as mericarp
Morphology Reproductive morphology Seeds Endosperm
Endosperm absent.
Distribution
Nine species, one in Vietnam, the others in Madagascar and eastern and southern Africa.

[FZ]

Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

Morphology General Habit
Trees or shrubs, unarmed or with small axillary spines; stems sometimes subscandent Trees or shrubs, unarmed or with small axillary spines; stems sometimes subscandent.
Morphology Leaves
Leaves deciduous, petiolate, opposite, simple, entire dentate or sublobed Leaves deciduous, petiolate, opposite, simple, entire dentate or sublobed.
Morphology Reproductive morphology Inflorescences
Inflorescences cymose, axillary or crowded at apices of shoots and appearing terminal Inflorescences cymose, axillary or crowded at apices of shoots and appearing terminal.
Morphology Reproductive morphology Flowers Calyx
Calyx often coloured; tube shortly funnel-shaped; limb broadly expanded rotate or patelliform, shallowly to ± deeply 5-lobed, very accrescent and papyraceous with a ± raised reticulation in fruit Calyx often coloured; tube shortly funnel-shaped; limb broadly expanded rotate or patelliform, shallowly to ± deeply 5-lobed, very accrescent and papyraceous with a ± raised reticulation in fruit.
Morphology Reproductive morphology Flowers Corolla
Corolla strongly zygomorphic; tube usually rather short, gibbous, contracted below the limb (mainly in bud), split dorsally to near base of tube giving a 1-lipped appearance; limb 5-lobed with the lobes all on anterior side arranged in a single lip comprising 2 separate posterior lobes, 2 lateral lobes and a larger median anterior labelliform lobe Corolla strongly zygomorphic; tube usually rather short, gibbous, contracted below the limb (mainly in bud), split dorsally to near base of tube giving a 1-lipped appearance; limb 5-lobed with the lobes all on anterior side arranged in a single lip comprising 2 separate posterior lobes, 2 lateral lobes and a larger median anterior labelliform lobe.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, didynamous, inserted near the mouth of corolla tube; filaments rolled into a circle in the bud and distinctly exserted from split in the corolla tube at anthesis, parallel, ascending then curving downwards, with a tuft of scales at the base; anthers with parallel or ± divergent thecae Stamens 4, didynamous, inserted near the mouth of corolla tube; filaments rolled into a circle in the bud and distinctly exserted from split in the corolla tube at anthesis, parallel, ascending then curving downwards, with a tuft of scales at the base; anthers with parallel or ± divergent thecae.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary hairy, 2 carpellate, each carpel divided into 2 loculi; ovules one per locule; style long, ± equalling and curved like the stamens; stigma short, 2-fid
Morphology Reproductive morphology Fruits
Fruit dry, hard, turbinate, obpyramidal or obconical, sometimes ± strongly 4-lobed above, rounded or ± flat at the apex, separating at maturity in 4 (?sometimes 2) trigonous mericarps. Fruit dry, hard, turbinate, obpyramidal or obconical, sometimes ± strongly 4-lobed above, rounded or ± flat at the apex, separating at maturity in 4 (?sometimes 2) trigonous mericarps.
Note
The Madagascan and African species were formerly placed in Holmskioldia , however, they differ from this genus mainly in the strongly zygomorphic corolla with a rather short, gibbous, fissured tube (in Holmskioldia the corolla tube is rather long, subcylindrical and not gibbous or dorsally split).  Karomia is further distinguished by its sub-1-lipped corolla limb; by its stamens inserted near the mouth of corolla tube (not in the middle and not prolonged below the insertion by raised pubescent lines, as in Holmskioldia ), and rolled into a ring in the bud but not so in Holmskioldia ; and by the hard dry fruit, not drupaceous.  The affinities of Karomia are closest to Rotheca Raf.
Distribution
A genus of nine species, one in Viet Nam the others in Madagascar and eastern and southern Africa.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary hairy, 2 carpellate, each carpel divided into 2 loculi; ovules one per locule; style long, ± equalling and curved like the stamens; stigma short, 2-fid.

[FTEA]

Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

Morphology General Habit
Shrubs or trees, glabrous or pubescent, unarmed or with small axillary spines; stems sometimes ± subscandent
Morphology Leaves
Leaves deciduous, petiolate, entire or toothed
Morphology Reproductive morphology Inflorescences
Inflorescences cymose, axillary or crowded at apices of shoots and appearing terminal; bracts mostly small, deciduous
Morphology Reproductive morphology Flowers Calyx
Calyx often brightly coloured; tube very short, urceolate; limb broadly expanded, very accrescent to form an entire or 5-lobed usually venose rotate plate-like structure in fruit
Morphology Reproductive morphology Flowers Corolla
Corolla-tube gibbous, very asymmetric, contracted beneath the limb, split dorsally almost to the base; limb 5-lobed and ± 1-lipped, very irregularly formed of 2 separated posterior lobes, 2 lateral lobes and a larger median anterior labelliform cucullate lobe
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens inserted near throat, the filaments enrolled into a ring in bud, distinctly exserted at anthesis and curved; filaments not prolonged beneath the point of insertion
Morphology Reproductive morphology Flowers Gynoecium Style
Style enrolled in a ring with the filaments in bud
Morphology Reproductive morphology Fruits
Fruit dry and hard, turbinate or turbinate-obpyramidal, rounded above or ± flat and with 4 small horns in a cross, at length dividing into ?2–4 pyrenes
Figures
Fig. 11
Distribution
Nine species, all but one formerly placed in Holmskioldia, occurring in Vietnam, E., SE. and S. Africa and particularly Madagascar
Note
Apart from one species occasionally cultivated in East Africa a recently described but imperfectly known endemic species may belong to this genus. R. Fernandes has given good reasons for recognising Karomia; she places the Madagascan and Vietnam species in sect. Karomia and the African species in sect. Cyclonemoides R. Fernandes. K. gigas had not then been described although she was aware of its existence and thought it unlikely to be a Holmskioldia.

Native to:

Kenya, KwaZulu-Natal, Madagascar, Malawi, Mozambique, Northern Provinces, Swaziland, Tanzania, Vietnam, Zambia, Zimbabwe

Karomia Dop appears in other Kew resources:

First published in Bull. Mus. Natl. Hist. Nat., sér. 2, 4: 1052 (1932)

Accepted by

  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

Literature

Flora Zambesiaca

  • Bull. Mus. Hist. Nat. (Paris), sér. 2, 4 : 1052 (1932).
  • Garcia de Orta, Sér. Bot. 7 : 36 (1988).
  • Harley et al. in Kadereit (ed.), Fam. Gen. Vasc. Pl. (Kubitzki, ed. in chief) VII: 198 (2004).
  • Kadereit (ed.), Fam. Gen. Vasc. Pl. (Kubitzki, ed. in chief) VII : 198 (2004).
  • R. Fernandes in Garcia de Orta, Sér. Bot. 7: 36 (1988).
  • in Bull. Mus. Hist. Nat. (Paris), sér. 2, 4: 1052 (1932).

Flora of Tropical East Africa

  • R. Fernandes in Garcia de Orta, sér. Bot. 7: 36 (1988)
  • in Bull. Mus. Hist. Nat. Paris, sér. 2, 4: 1052 (1932)

Lamiaceae Key Genus Fact Sheets

  • Harley RM, Atkins S, Budantsev AL, Cantino PD, Conn BJ, Grayer R, Harley MM, de Kok RPJ, Krestovskaja T, Morales R, Paton AJ, Ryding O, and Upson T. 2004. Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Sp
  • Karomia Dop, Bull. Mus. Hist. Hat. (Paris), ser. 2, 4: 1052 (1932)
  • R. Fern., Garcia de Orta, Ser. Bot., Lisboa 7: 33-46 (1985).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Lamiaceae Key Genus Fact Sheets
Nina Davies, Gemma Bramley and Don Kirkup, Royal Botanic Gardens, Kew
http://creativecommons.org/licenses/by/3.0