1. Family: Rutaceae Juss.
    1. Vepris Comm. ex A Juss.

      1. This genus is accepted, and its native range is Africa, India.

    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Trees or shrubs, sometimes subscandent.
    Leaves
    Leaves alternate or rarely opposite, 1- or 3-foliolate; leaflets dotted with pellucid glands.
    Inflorescences
    Inflorescence of axillary and terminal racemes or panicles or clustered cymes, sometimes on older leafless branches.
    Flowers
    Flowers (2–3) 4-merous, unisexual by abortion. Male flowers: stamens (6–7) 8, inserted at the base of the disk, those opposite the petals shorter than those opposite the sepals; filaments flattened at the base, subulate; anthers dorsifixed, 2-lobed at the base; ovary vestigial with 2–4 styles. Female flowers: staminodes 8 or fewer; ovary (1) 2–4-locular; loculi 2-ovulate; ovules pendulous; stigma sessile, 2–4-lobed.
    Calyx
    Sepals united to form a cupuliform (3) 4-lobed calyx.
    Corolla
    Petals (3) 4, oblong, imbricate in bud.
    Fruits
    Fruit drupaceous, (1) 2–4-locular; loculi 1-seeded.
    Seeds
    Endosperm present.
    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Unarmed shrubs or trees, sometimes subscandent
    Leaves
    Leaves 1–3-foliolate, alternate or rarely opposite; leaflets dotted with pellucid glands
    Inflorescences
    Inflorescence of axillary and terminal racemes, panicles or clustered cymes
    Flowers
    Flowers bisexual on unisexual by abortion, (2–3–)4-merous Male flowers: stamens (6–7–)8(–9), very rarely 5, inserted at the base of a disk, those opposite the sepals longer than the ones opposite the petals; filaments flattened at the base, subulate; anthers dorsifixed, 2-lobed at the base; ovary vestigial with 2–4 styles Female flowers: staminodes 8 or fewer; ovary (1–)2–4-locular; each locule with (I–)2 pendulous ovules; stigma capitate or 2–4-lobed, sessile
    Calyx
    Sepals united to form a cupuliform (3–)4-lobed calyx
    Corolla
    Petals (3–)4, oblong, imbricate in bud
    Male
    Male flowers: stamens (6–7–)8(–9), very rarely 5, inserted at the base of a disk, those opposite the sepals longer than the ones opposite the petals; filaments flattened at the base, subulate; anthers dorsifixed, 2-lobed at the base; ovary vestigial with 2–4 styles
    Female
    Female flowers: staminodes 8 or fewer; ovary (1–)2–4-locular; each locule with (I–)2 pendulous ovules; stigma capitate or 2–4-lobed, sessile
    Fruits
    Fruit drupaceous, (1–)2–4-locular; locule 1(–2)-seeded.
    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Trees or shrubs.
    Leaves
    Leaves alternate, ( 1 ) 3-foliolate; leaflets densely dotted with pellucid glands.
    Inflorescences
    Inflorescence of axillary and terminal panicles, racemes or cymose clusters.
    Calyx
    Sepals united into a 4–5-lobed cupuliform calyx; lobes short, imbricate.
    Corolla
    Petals longer than the calyx, imbricate in bud.
    Flowers
    Male flowers: stamens 4–5 inserted at the base of the disk; filaments linear, subulate; anthers dorsifixed, cordate, deeply 2-lobed at the base; ovary-vestigial. Female flowers: staminodes 4–5 or fewer, opposite the sepals; ovary 1-locular with 2 pendulous ovules; style short; stigma broadly peltate. Flowers 4–5-merous, unisexual by abortion, rarely bisexual.
    Fruits
    Fruit drupaceous, globose or ovoid or ellipsoid, 1-seeded; endocarp woody.
    Seeds
    Seed ovoid, testa thin; endosperm absent.
    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Unarmed shrubs or trees
    Leaves
    Leaves 1–3-foliolate; leaflets densely dotted with pellucid glands
    Inflorescences
    Inflorescence of axillary and terminal panicles, racemes or cymose clusters
    Flowers
    Flowers unisexual by abortion, rarely bisexual, 4–5-merous Female flowers: staminodes 4–5 or fewer; ovary unilocular with 2 pendulous ovules; style short; stigma broadly peltate, entire or lobed Male flowers: stamens 4–5, inserted at the base of the disk; filaments linear, subulate; anthers dorsifixed or basifixed, deeply 2-lobed at the base; ovary vestigial
    Calyx
    Sepals united into a 4–5-lobed cupuliform calyx; lobes short and imbricate
    Corolla
    Petals longer than the calyx, imbricate in bud
    Male
    Male flowers: stamens 4–5, inserted at the base of the disk; filaments linear, subulate; anthers dorsifixed or basifixed, deeply 2-lobed at the base; ovary vestigial
    Female
    Female flowers: staminodes 4–5 or fewer; ovary unilocular with 2 pendulous ovules; style short; stigma broadly peltate, entire or lobed
    Fruits
    Fruit drupaceous, globose to oblong-ovoid, 1-seeded; endocarp woody
    Seeds
    Seed ovoid; testa thin.
    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Shrubs or small trees.
    Leaves
    Leaves alternate, 3-foliolate; leaflets densely dotted with pellucid glands.
    Inflorescences
    Inflorescence of short racemes or panicles, axillary and terminal.
    Flowers
    Flowers unisexual by abortion. Male flowers: stamens 8, inserted at the base of the disk; filaments subulate; anthers cordate; ovary vestigial. Female flowers: staminodes 8, filiform, with atrophied anthers; ovary slightly 4-lobed with thick verrucose walls, 4-locular; loculi 2-ovulate; stigma sessile, 4-lobed.
    Calyx
    Sepals 4, ovate, united at the base.
    Fruits
    Fruit very woody, globose, subtetragonal, 4-locular; loculi 1-seeded.
    Seeds
    Seed oblong; endosperm absent.
    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Shrubs or small trees
    Leaves
    Leaves trifoliolate, alternate; leaflets oblanceolate or obovate, densely dotted with pellucid glands
    Inflorescences
    Inflorescence of short racemes or panicles, axillary or terminal
    Flowers
    Flowers unisexual by abortion Male flowers: stamens 8, inserted at the base of the disk; filaments subulate; anthers cordate; ovary vestigial Female flowers: ovary slightly 4-lobed with thick verrucose walls, 4-locular; locules 2-ovulate; stigma sessile, 4-lobed; staminodes 8, filiform
    Calyx
    Sepals 4, ovate, connate at the base
    Male
    Male flowers: stamens 8, inserted at the base of the disk; filaments subulate; anthers cordate; ovary vestigial
    Female
    Female flowers: ovary slightly 4-lobed with thick verrucose walls, 4-locular; locules 2-ovulate; stigma sessile, 4-lobed; staminodes 8, filiform
    Fruits
    Fruit a woody nut, globose, tetragonal or subtetragonal, verrucose, 4-locular; locules 1-seeded
    Seeds
    Seeds oblong.
    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Trees with alternate or opposite 3–5-foliolate leaves.
    Inflorescences
    Inflorescence of terminal and axillary panicles.
    Flowers
    Flowers 4-merous, unisexual by abortion. Male flowers: stamens 4, opposite the sepals; filaments short; anthers introrse; carpels vestigial. Female flowers: staminodes 4; carpels 2–4 (1–3 aborting), slightly united at the base and by the stigma (otherwise free) into a 2–4-locular ovary with 2 pendulous apical ovules in each loculus; style short; stigma discoid, 2–4-lobed.
    Calyx
    Sepals 4.
    Corolla
    Petals 4, valvate or slightly imbricate.
    Fruits
    Fruit of 1–4 1-seeded drupes.
    Seeds
    Seed with a thin testa; endosperm absent.
    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Shrubs or small unarmed trees
    Leaves
    Leaves trifoliolate; leaflets sessile to shortly petiolulate, glandular-punctate
    Inflorescences
    Inflorescence of terminal and axillary panicles
    Flowers
    Flowers polygamous
    Calyx
    Calyx with 4 shallow lobes
    Corolla
    Petals 4
    Stamens
    Stamens 4
    Disc
    Disk with long stiff yellow hairs
    Pistil
    Ovary glabrous or sparingly hispid; carpels 2, united; styles 1–2; stigma peltate; ovules 2
    Fruits
    Fruit 2-lobed, occasionally one carpel aborting and forming a lobe at the base
    Seeds
    Seeds solitary in each carpel.
    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Shrubs with pilose branches, petioles and inflorescences.
    Leaves
    Leaves alternate, 3-foliolate; petiole subterete; leaflets sessile, elliptic or obovate, densely dotted with minute pellucid glands; nerves few, remote from each other and anastomosing in arches near the margin.
    Inflorescences
    Inflorescence of terminal and axillary panicles; bracts and bracteoles small, caducous.
    Calyx
    Sepals 4, slightly united at the base or free.
    Corolla
    Petals 4, valvate.
    Flowers
    Male flowers: stamens usually 8, inserted at the base of the disk; vestigial ovary of 2 free carpels with atrophied styles. Female flowers: staminodes 8, short or vestigial; ovary of 2 (3) free 1-locular carpels, each with 2 pendulous apical ovules; styles united at the apex in a discoid peltate stigma. Flowers 4-merous, unisexual by abortion, dioecious.
    Fruits
    Fruit unknown.
    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Shrubs or trees with alternate branches, dioecious
    Leaves
    Leaves alternate to nearly opposite, trifoliolate; leaflets sessile or shortly petiolulate, densely dotted with minute pellucid glands; nerves few, remote from each other and netted near the margin
    Inflorescences
    Inflorescence of terminal and axillary panicles; bracts and bracteoles small, caducous
    Flowers
    Flowers 4-merous, unisexual by abortion Male flowers: stamens 8, inserted at the base of the disk, vestigial ovary of 2 free carpels with atrophied styles Female flowers: staminodes 8, short or vestigial; ovary of 2(–3) free unilocular carpels, each with 2 pendulous apical ovules; styles united at the apex in a discoid peltate stigma
    Calyx
    Sepals 4, slightly united at the base or free
    Corolla
    Petals 4, ± valvate in bud
    Male
    Male flowers: stamens 8, inserted at the base of the disk, vestigial ovary of 2 free carpels with atrophied styles
    Female
    Female flowers: staminodes 8, short or vestigial; ovary of 2(–3) free unilocular carpels, each with 2 pendulous apical ovules; styles united at the apex in a discoid peltate stigma
    Fruits
    Fruits usually paired, fleshy, ellipsoid and 1-seeded.

    Images

    Distribution

    Native to:

    Angola, Benin, Botswana, Burkina, Burundi, Cameroon, Cape Provinces, Caprivi Strip, Central African Repu, Comoros, Congo, Djibouti, Eritrea, Ethiopia, Gabon, Ghana, Guinea, Gulf of Guinea Is., India, Ivory Coast, Kenya, KwaZulu-Natal, Liberia, Madagascar, Malawi, Mali, Mauritius, Mozambique, Niger, Nigeria, Northern Provinces, Rodrigues, Rwanda, Réunion, Saudi Arabia, Sierra Leone, Somalia, Sudan, Swaziland, Tanzania, Uganda, Yemen, Zambia, Zaïre, Zimbabwe

    Vepris Comm. ex A Juss. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Dec 1, 2003 Pollard, B.J. [930], Cameroon 76534.000
    Richard, A. [221], Madagascar 33526.000
    Dowsett-Lemaire, F. [2529], Mozambique K000614139
    Harris, T. [602], Mozambique K000614191
    Patel, H. [7211], Mozambique K000545172
    Patel, H. [7214], Mozambique K000545175
    Timberlake, J. [5398], Mozambique K000614390
    Mphamba [56], Mozambique K000614275
    Olorunfemi [30709], Cameroon Oricia 21868.000

    First published in Mém. Mus. Hist. Nat. 12: 509 (1825)

    Literature

    Flora Zambesiaca
    • in Mem. Mus. Hist. Nat. Par. 12: 509 (1825).
    Flora of Tropical East Africa
    • in Mém. Mus. Nat. Hist. Nat. Paris 12: 509 (1825)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0