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This genus is accepted, and its native range is America.
Physalis sp.

[FTEA]

Solanaceae, Jennifer M Edmonds. Oliganthes, Melongena & Monodolichopus, Maria S. Vorontsova & Sandra Knapp. Flora of Tropical East Africa. 2012

Morphology General Habit
Annual or perennial herbs, usually much-branched, sometimes woody basally.
Morphology Stem
Stems sometimes viscid, hairs simple, branched or stellate, glandular or eglandular
Morphology Leaves
Leaves alternate, rarely opposite
Morphology Reproductive morphology Inflorescences
Inflorescences axillary, usually composed of a single flower, occasionally 2–4(–7) flowered fascicles; flowers erect to pendent, hermaphrodite
Morphology Reproductive morphology Flowers Calyx
Calyx campanulate to cupulate, actinomorphic with 5 lobes or 5-partite with short lobes; enlarged and persistent in fruit when basally invaginated
Morphology Reproductive morphology Flowers Corolla
Corolla broadly campanulate to rotate or funnelform, rarely urceolate, usually actinomorphic, often with dark basal spots and densely pubescent inner ring alternating with stamens internally, margin entire or tube terminating in 5 short lobes
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 5, usually equal and exserted; filaments fused to lower part of corolla tube where broader, glabrous or sparsely pilose; anthers equal or unequal, oblong, bilobed, basi- or dorsi-fixed, sometimes convergent around stigma
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary superior, bilocular, ovules numerous, placentation axile; disc annular, occasionally absent; style filiform, usually glabrous and exserted; stigma discoid-capitate, bilobed
Morphology Reproductive morphology Fruits
Fruit a berry, mature pericarp thin and translucent, enclosed by enlarged and inflated accrescent chartaceous usually reticulately-veined bladder-like urceolate calyx which 5-angled or prominently 10-ribbed, sometimes with 5 basal auricles, often brightly coloured, the mouth usually almost completely closed by connivent calyx lobes or teeth
Morphology Reproductive morphology Seeds
Seeds numerous, vesicular to foveolate, sclerotic granules absent.
Note
A few species are cultivated world-wide for their ornamental showy calyces, or for their edible berries and they are collectively known as Husk-tomatoes or Ground-cherries. Hepper (in Fl. Ceylon, 4: 391 (1987)) described Physalis as being an extremely puzzling genus of considerable taxonomic and nomenclatural complexity, while Symon (in Journ. Adelaide Bot. Gard. 3(2): 149 (1981)) thought that many of the nomenclatural problems would remain insoluble until African and Asian names and taxa were studied and compared with those in America. Older literature contains numerous synonyms for each of the Physalis species found in the floral region; where the names concerned have not been encountered on any East African material, they have not been included in this treatment. With the exception of P. peruviana, it is often difficult to determine which species is actually being described in many of the Floras cited, and the associated nomenclature is extremely complex. It is clearly a genus in urgent need of worldwide revision.

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Herbs, glabrous or with indumentum of simple hairs
Morphology Leaves
Leaves alternate or paired, simple
Morphology Reproductive morphology Flowers
Flowers solitary, axillary
Morphology Reproductive morphology Flowers Calyx
Calyx campanulate, lobed, much enlarged in fruit
Morphology Reproductive morphology Flowers Corolla
Corolla campanulate to rotate, subentire or lobed
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens included, inserted near base of corolla-tube; anthers dehiscing longitudinally
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 2-celled, with numerous ovules; style slender, stigma capitate
Morphology Reproductive morphology Fruits
Fruit a globose berry surrounded by the enlarged calyx
Morphology Reproductive morphology Seeds
Seeds flattened, minutely pitted.
Distribution
Genus of some 100 species, the large majority in America.

[FZ]

Flora Zambesiaca. Vol. 8, Part 4. Solanaceae. Gonçalves AE. 2005

Morphology General
Herbs or more rarely shrubs, sometimes sarmentose, glabrous to hairy; indumentum varied and often intermixed, the hairs multicellular (“jointed”) and short, unicellular, simple or occasionally stellate, often glandular or viscid, more rarely branched, also stipitate or sessile glands; rhizomatous structures sometimes present
Morphology Leaves
Leaves solitary, alternate, sometimes 2 or 3 appearing together, one larger than the others, petiolate, entire to coarsely lobed, rarely pinnatifid Leaves solitary, alternate, sometimes 2 or 3 appearing together, one larger than the others, petiolate, entire to coarsely lobed, rarely pinnatifid.
Morphology Reproductive morphology Flowers
Flowers solitary, extra-axillary or appearing axillary (elsewhere occasionally up to several flowers in small, axillary fascicles or rarely in terminal pseudoracemes), actinomorphic. Flowers solitary, extra-axillary or appearing axillary (elsewhere occasionally up to several flowers in small, axillary fascicles or rarely in terminal pseudoracemes), actinomorphic; pedicel slender, often elongated and drooping or nodding in fruit; bracts obsolete.
Morphology Reproductive morphology Flowers Pedicel
Pedicel slender, often elongated and drooping or nodding in fruit; bracts obsolete
Morphology Reproductive morphology Flowers Calyx
Calyx often as long as the corolla tube, campanulate or sometimes tubular-campanulate, 5-lobed to sub-truncate, the lobes if present as long as or shorter than the tube, often splitting at the sutures, conniving at the apex, with valvate aestivation; in fruit much enlarged and usually bladdery-inflated, often splitting at the sutures, loosely enclosing it (or elsewhere nearly so), 5- or 10-angled to -ribbed, sometimes ± terete, reticulate, often invaginated basally, contracted apically with a small opening surrounded by the lobes, remaining membranous or becoming chartaceous Calyx often as long as the corolla tube, campanulate or sometimes tubular-campanulate, 5-lobed to sub-truncate, the lobes if present as long as or shorter than the tube, often splitting at the sutures, conniving at the apex, with valvate aestivation; in fruit much enlarged and usually bladdery-inflated, often splitting at the sutures, loosely enclosing it (or elsewhere nearly so), 5- or 10-angled to -ribbed, sometimes ± terete, reticulate, often invaginated basally, contracted apically with a small opening surrounded by the lobes, remaining membranous or becoming chartaceous.
Morphology Reproductive morphology Flowers Corolla
Corolla yellowish to whitish, often blotched with 5 darker markings, campanulate, rarely infundibular-rotate (or elsewhere shortly infundibular); tube short, ± hairy inside above insertion of stamens and ± glabrous; limb plicate, subentire, sinuate, 5-angled or shortly and widely 5-lobed (to exceptionally 5-parted elsewhere), sometimes ± reflexed; lobes ovate-deltate (to subulate elsewhere), with imbricate-quincuncial aestivation Corolla yellowish to whitish, often blotched with 5 darker markings, campanulate, rarely infundibular-rotate (or elsewhere shortly infundibular); tube short, ± hairy inside above insertion of stamens and ± glabrous; limb plicate, subentire, sinuate, 5-angled or shortly and widely 5-lobed (to exceptionally 5-parted elsewhere), sometimes ± reflexed; lobes ovate-deltate (to subulate elsewhere), with imbricate-quincuncial aestivation.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 5, mostly ± equal, usually inserted near the base of the corolla tube, ± included or slightly exserted; filaments slender (to nearly as wide as the anthers elsewhere), often ± straight, pubescent or glabrous; anthers oblong or linear-oblong to ovate in outline, straight or occasionally twisting and curling after anthesis, free or sometimes conniving, attached on the lower part of the back or at the base between the thecae, dehiscing by longitudinal slits Stamens 5, mostly ± equal, usually inserted near the base of the corolla tube, ± included or slightly exserted; filaments slender (to nearly as wide as the anthers elsewhere), often ± straight, pubescent or glabrous; anthers oblong or linear-oblong to ovate in outline, straight or occasionally twisting and curling after anthesis, free or sometimes conniving, attached on the lower part of the back or at the base between the thecae, dehiscing by longitudinal slits.
Morphology Reproductive morphology Flowers Disc
Disk annular, adnate to and surrounding the basal part of the ovary, to none Disk annular, adnate to and surrounding the basal part of the ovary, to none.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 2-locular; ovules hemicampylotropous, few–numerous in each locule on a much enlarged, subglobose placenta adnate to the dissepiment near the axis.
Morphology Reproductive morphology Flowers Gynoecium Style
Style slender to stout.
Morphology Reproductive morphology Flowers Gynoecium Stigma
Stigma capitate to nearly truncate, sometimes 2-lobed to -lamellate, included or exserted
Distribution
Genus with c. 90 species, mostly centred in the warm regions of America mainly in Mexico; widely distributed in warm and temperate regions of the Old World and Australia, with several species introduced and a few widespread as weeds or in cultivation; 7 s
Morphology General Habit
Herbs or more rarely shrubs, sometimes sarmentose, glabrous to hairy; indumentum varied and often intermixed, the hairs multicellular (“jointed”) and short, unicellular, simple or occasionally stellate, often glandular or viscid, more rarely branched, also stipitate or sessile glands; rhizomatous structures sometimes present.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary 2-locular; ovules hemicampylotropous, few–numerous in each locule on a much enlarged, subglobose placenta adnate to the dissepiment near the axis; style slender to stout; stigma capitate to nearly truncate, sometimes 2-lobed to -lamellate, included or exserted.
Morphology Reproductive morphology Fruits
Fruit baccaceous, mostly ± globose, sessile or sometimes on a short gynobase, juicy or fleshy, often thin-walled, sometimes viscid, 2-locular. Fruit baccaceous, mostly ± globose, sessile or sometimes on a short gynobase, juicy or fleshy, often thin-walled, sometimes viscid, 2-locular
Morphology Reproductive morphology Seeds
Seeds few–numerous, flattened, laterally compressed, suborbicular to reniform; testa somewhat leathery, slightly rugose or smooth, glabrous; embryo ± circinnate, flattened, subperipheral, in a fleshy or horny endosperm, with a terete radicle as wide as the semi-terete cotyledons. Seeds few–numerous, flattened, laterally compressed, suborbicular to reniform; testa somewhat leathery, slightly rugose or smooth, glabrous; embryo ± circinnate, flattened, subperipheral, in a fleshy or horny endosperm, with a terete radicle as wide as the semi-terete cotyledons

Native to:

Afghanistan, Alabama, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Aruba, Assam, Bahamas, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, California, Cayman Is., Chile Central, Chile North, China North-Central, Colombia, Colorado, Costa Rica, Cuba, Dominican Republic, East Himalaya, Ecuador, El Salvador, Florida, French Guiana, Galápagos, Georgia, Guatemala, Guyana, Haiti, Honduras, Idaho, Illinois, India, Indiana, Iowa, Jamaica, Kansas, Kentucky, Leeward Is., Louisiana, Manitoba, Mexican Pacific Is., Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Minnesota, Mississippi, Missouri, Montana, Nebraska, Nepal, Netherlands Antilles, Nevada, New Jersey, New Mexico, New York, Nicaragua, North Carolina, North Dakota, Ohio, Oklahoma, Ontario, Oregon, Panamá, Paraguay, Pennsylvania, Peru, Puerto Rico, Québec, South Carolina, South Dakota, Southwest Caribbean, Suriname, Tennessee, Texas, Trinidad-Tobago, Turks-Caicos Is., Uruguay, Utah, Venezuela, Venezuelan Antilles, Vermont, Virginia, Washington, West Virginia, Windward Is., Wisconsin, Wyoming

Introduced into:

Algeria, Andaman Is., Angola, Ascension, Austria, Azores, Baleares, Bangladesh, Belgium, Benin, Bermuda, Bismarck Archipelago, Borneo, Botswana, Bulgaria, Burkina, Burundi, Cambodia, Cameroon, Canary Is., Cape Provinces, Cape Verde, Caroline Is., Central African Repu, Central European Rus, Chad, China South-Central, China Southeast, Christmas I., Cocos (Keeling) Is., Comoros, Congo, Cook Is., Czechoslovakia, Djibouti, Easter Is., Egypt, Equatorial Guinea, Eritrea, Ethiopia, Fiji, Free State, Gabon, Gambia, Ghana, Gilbert Is., Great Britain, Greece, Guinea, Guinea-Bissau, Gulf of Guinea Is., Gulf States, Hainan, Hawaii, Hungary, Italy, Ivory Coast, Japan, Jawa, Juan Fernández Is., Kazan-retto, Kenya, Kirgizstan, Korea, KwaZulu-Natal, Laccadive Is., Laos, Lebanon-Syria, Lesotho, Lesser Sunda Is., Liberia, Libya, Line Is., Madagascar, Madeira, Malawi, Malaya, Maldives, Mali, Manchuria, Marianas, Marquesas, Marshall Is., Maryland, Mauritania, Mauritius, Morocco, Mozambique, Myanmar, Namibia, Nauru, New Caledonia, New Guinea, New South Wales, New Zealand North, Nicobar Is., Niger, Nigeria, Niue, Norfolk Is., Northern Provinces, Northern Territory, Ogasawara-shoto, Oman, Pakistan, Palestine, Philippines, Phoenix Is., Portugal, Primorye, Queensland, Romania, Rwanda, Réunion, Samoa, Saudi Arabia, Senegal, Seychelles, Sierra Leone, Society Is., Socotra, Solomon Is., Somalia, South Australia, South China Sea, Spain, Sri Lanka, St.Helena, Sudan, Swaziland, Tadzhikistan, Taiwan, Tanzania, Tasmania, Thailand, Togo, Tonga, Tristan da Cunha, Tuamotu, Tubuai Is., Turkey, Tuvalu, Uganda, Ukraine, Uzbekistan, Victoria, Vietnam, Wallis-Futuna Is., West Himalaya, West Siberia, Western Australia, Yemen, Yugoslavia, Zambia, Zaïre, Zimbabwe

Physalis L. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Willis, F.R. [123], New Guinea 70592.000
Hinton, G.B. [8174], Mexico K000042324
Hinton, G.B. [8200], Mexico K000042325
Hinton, G.B. [8457], Mexico K000042327
Palmer, E. [165], Mexico K000042328
Palmer, E. [622], Mexico K000042329
Palmer, E. [634], Mexico K000042332
Palmer, E. [463], Mexico K000042337
Nelson, E.W. [4649], Mexico K000042338
Rose, J.N. [7172], Mexico K000042339
Balls, E.K. [B5413], Mexico K000042341
Palmer, E. [144], Mexico K000042345
Palmer, E. [220], Mexico K000042349
Palmer, E. [100], Mexico K000042350
Hinton, G.B. [4202], Mexico K000042354
Hinton, G.B. [4686], Mexico K000042355
Hinton, G.B. [6347], Mexico K000042359
Hinton, G.B. [8472], Mexico K000042363
Botteri, M. [853], Mexico K000042368
Coulter, T. [221], Mexico K000042372
Muller, N. [1336], Mexico K000042373
Palmer, E. [53], Mexico K000042351
Schaffner, J.G. [699], Mexico K000042348
Hinton, G.B. [13156], Mexico K000042344
Hinton, G.B. [6347], Mexico K000042370
Rico, L. [1411], Bolivia K000295481
s.coll. [Cat. no. s.n.], Myanmar K001132461
Finlayson, G. [Cat. no. s.n.] K001132465
s.coll. [Cat. no. s.n.] K001132466
Palmer, E. [480], Mexico K000042366
Palmer, E. [379], Mexico K000042347
Hinton, G.B. [4686], Mexico K000042367
Hinton, G.B. [5823], Mexico K000042360
Palmer, E. [494], Mexico K000042333
Palmer, E. [297], Mexico K000042331
Balls, E.K. [B4404], Mexico K000042342
s.coll. [Cat. no. s.n.] K001132965
s.coll. [Cat. no. s.n.] K001132460
s.coll. [Cat. no. s.n.] K001132458
Heyne, B. [Cat. no. s.n.] K001132459
Balls, E.K. [B4714], Mexico K000042340
Bourgeau, M. [3228], Mexico K000042362
Bourgeau, M. [2940], Mexico K000042371
Orcutt, C.R. [3531], Mexico K000042385
Hinton, G.B. [8198], Mexico K000042364
Hinton, G.B. [6687], Mexico K000042356
s.coll. [Cat. no. s.n.], Myanmar K001132464
Palmer, E. [463], Mexico K000042334
Matuda, E. [3312], Mexico K000042335
s.coll. [Cat. no. s.n.] K001132463
Rovirosa, J.N. [558], Mexico K000042386
Hinton, G.B. [8034], Mexico K000042365
s.coll. [Cat. no. s.n.] K001132457
Balls, E.K. [4331], Mexico K000042285
s.coll. [Cat. no. s.n.] K001132462
Trail, J.W.H. [601], Brazil K001071980
Langlassé, E. [309], Mexico K000042352
Nelson, E.W. [6272], Mexico K000042336
Balls, E.K. [B4715], Mexico K000042343
Hinton, G.B. [11928], Mexico K000042326
Palmer, E. [127], Mexico K000042346
Hinton, G.B. [619], Mexico K000042353
Palmer, E. [621], Mexico K000042330
Hinton, G.B. [8606], Mexico Margaranthus K000063226

First published in Sp. Pl.: 182 (1753)

Accepted by

  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.

Literature

Flora of West Tropical Africa

  • O. E. Schulz in Urban, Symb. Antill. 6: 140 (1909)
  • Waterfall in Rhodora 60: 107—114, 128—142, 152—173 (1958).
  • —F.T.A. 4, 2: 246

Flora Zambesiaca

  • Gen. Pl., ed. 5: 85 (1754).
  • Hunziker, Gen. Solanacearum: 202 (2001).
  • Sp. Pl.: 182 (1753)

Flora of Somalia

  • Flora Somalia, Vol 3, (2006) Author: by M. Thulin [updated by M. Thulin 2008]

Flora of Tropical East Africa

  • DC., Prodr. 13(1):434 (1852);
  • Gen. Pl. ed. 5: 85 (1754);
  • Gen. Solanaceae: 202–208 (2001)
  • Rhodora 60: 107–114, 128–142, 152–173 (1958)
  • Rhodora 69: 82–120, 203–239, 319–329 (1967);
  • Sp. Pl. 1: 182 (1753)

Flora Zambesiaca
Flora Zambesiaca
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Flora of Somalia
Flora of Somalia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Science Photographs
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/