Skip to main content
This species is accepted, and its native range is Central & S. U.S.A. to Tropical & Subtropical America.
Cenchrus echinatus L.

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Nativa en Colombia; Alt. 0 - 1500 m.; Andes, Guayana y Serranía de La Macarena, Islas Caribeñas, Llanura del Caribe, Orinoquia, Sierra Nevada de Santa Marta, Valle del Cauca, Valle del Magdalena.
Morphology General Habit
Hierba
Conservation
No Evaluada

[FTEA]

Gramineae, W. D. Clayton, S. M. Phillips & S. A. Renvoize. Flora of Tropical East Africa. 1974

Morphology General Habit
Coarse annual; culms 15–90 cm. high, ascending.
Morphology Leaves
Leaf-blades 4–25 cm. long, 3–10 mm. wide.
Morphology Reproductive morphology Inflorescences
Panicle 2–10 cm. long; involucres globose, 5–10 mm. long; inner spines flattened, connate for ± 1/2 their length to form a deep cup, pubescent, retrorsely barbellate; outer spines rigid, mostly shorter than the inner, ± divergent.
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets 2–3 per burr, 5–7 mm. long.
Habitat
Roadside and waste places; 0–900 m.
Distribution
now established as a weed throughout the tropics and subtropics K7 T3 T6

[UNAL]
Vernacular
Pega-pega, Purula

[FWTA]

Gramineae, W. D. Clayton. Flora of West Tropical Africa 3:2. 1972

Morphology General Habit
Annual 30–90 cm. high.

[GB]
Morphology General Habit
Annual. Culms geniculately ascending; 15-90 cm long. Ligule a fringe of hairs. Leaf-blades 4-25 cm long; 3-10 mm wide.
Morphology Reproductive morphology Inflorescences
Inflorescence a panicle. Panicle spiciform; linear; interrupted (or lax); 2-10 cm long. Primary panicle branches accrescent to a central axis; with sessile scars on axis. Panicle axis angular; bearing deciduous spikelet clusters. Spikelets subtended by an involucre. Fertile spikelets sessile; 2-3 in the cluster. Involucre composed of bristles; connate into a cup below; with 2-5 mm connate; globose; 5-10 mm long; base obconical. Involucral bristles deciduous with the fertile spikelets; with an outer whorl of thinner bristles; inner bristles longer than outer; with longest bristle scarcely emergent; 2-5 mm long; flattened; rigid; retrorsely scaberulous; pubescent; spinose.
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets ovate; dorsally compressed; acuminate; 5-7 mm long; falling entire; deciduous with accessory branch structures.
Fertile
Spikelets comprising 1 basal sterile florets; 1 fertile florets; without rhachilla extension. Spikelets ovate; dorsally compressed; acuminate; 5-7 mm long; falling entire; deciduous with accessory branch structures.
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes shorter than spikelet; thinner than fertile lemma. Lower glume ovate; 1.3-3.4 mm long; 0.33-0.5 length of spikelet; membranous; without keels; 1 -veined. Lower glume lateral veins absent. Lower glume apex obtuse, or acute. Upper glume ovate; 3.8-5.7 mm long; 0.75 length of spikelet; membranous; without keels; 3-5 -veined. Upper glume surface puberulous. Upper glume apex obtuse, or acute.
Morphology Reproductive morphology Flowers Florets
Basal sterile florets barren; with palea. Lemma of lower sterile floret ovate; 1 length of spikelet; membranous; 5 -veined; obtuse, or acute. Fertile lemma ovate; 5-7 mm long; coriaceous; much thinner on margins; without keel; 5 -veined. Lemma margins flat. Lemma apex obtuse. Palea coriaceous.
Morphology Reproductive morphology Fruits
Caryopsis with adherent pericarp; ellipsoid; dorsally compressed; 3 mm long; truncate.
Distribution
Africa: north, Macaronesia, west tropical, west-central tropical, east tropical, and western Indian ocean. Asia-temperate: western Asia, Arabia, China, and eastern Asia. Asia-tropical: India, Indo-China, Malesia, and Papuasia. Australasia: Australia. Pacific: southwestern, south-central, northwestern, and north-central. North America: southwest USA, south-central USA, southeast USA, and Mexico. South America: Mesoamericana, Caribbean, northern South America, western South America, Brazil, and southern South America.
Reference
Paniceae. FTEA.
Diagnostic
Involucre base 2.2-3.6mm wide.

Native to:

Alabama, Argentina Northeast, Argentina Northwest, Arizona, Arkansas, Aruba, Bahamas, Belize, Bermuda, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, California, Cayman Is., Central American Pac, Chile North, Chile South, Colombia, Costa Rica, Cuba, Desventurados Is., District of Columbia, Dominican Republic, Ecuador, El Salvador, Florida, French Guiana, Galápagos, Georgia, Guatemala, Guyana, Haiti, Honduras, Jamaica, Kansas, Kentucky, Leeward Is., Louisiana, Maryland, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mississippi, Netherlands Antilles, Nevada, New Mexico, Nicaragua, North Carolina, Oklahoma, Panamá, Paraguay, Peru, Puerto Rico, South Carolina, Southwest Caribbean, Suriname, Tennessee, Texas, Trinidad-Tobago, Turks-Caicos Is., Uruguay, Venezuela, Venezuelan Antilles, Virginia, Windward Is.

Introduced into:

Aldabra, Bangladesh, Borneo, Canary Is., Cape Verde, Caroline Is., Chagos Archipelago, China South-Central, China Southeast, Cocos (Keeling) Is., Cook Is., Czechoslovakia, Easter Is., Egypt, Equatorial Guinea, Fiji, Gabon, Ghana, Gilbert Is., Greece, Gulf States, Hainan, Hawaii, India, Ivory Coast, Japan, Kenya, Lebanon-Syria, Lesser Sunda Is., Line Is., Madagascar, Maine, Malaya, Marianas, Marquesas, Marshall Is., Massachusetts, Mauritius, Mexican Pacific Is., Mozambique Channel I, Nansei-shoto, Nauru, New Caledonia, New Guinea, Nigeria, Niue, Northern Territory, Oman, Palestine, Philippines, Phoenix Is., Rodrigues, Réunion, Samoa, Saudi Arabia, Seychelles, Society Is., Solomon Is., South China Sea, Sri Lanka, Taiwan, Tanzania, Togo, Tokelau-Manihiki, Tonga, Tuamotu, Tubuai Is., Tuvalu, Vanuatu, Vermont, Wake I., Wallis-Futuna Is.

Cenchrus echinatus L. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 1973 Anderson, W.R. [36950], Bahia K000004810
Jan 1, 1933 Curtis, A.H. [3619], Florida K001056122
Eiten, G. [4892], Bahia K000004803
Bolland, G. [1926], Brazil K000004804
Harley, R.M. [17273], Bahia K000004814
Gardner [1190], Pernambuco K000004805
Gardner [1190], Pernambuco K000004806
Bommer, D. [44], Brazil K000004807
Moseley [0], Fernando de Noronha K000004808
Mattos Silva, L.A. [3040], Sergipe K000004809
Santos, T.S. [3680], Bahia K000004811
Hage, J.L. [1582], Bahia K000004812
Mori, S.A. [11368], Bahia K000004813
Davidse, G. [11649], Bahia K000004815
Harley, R.M. [16243], Bahia K000004816
Grupo Pedra do Cavalo [62], Bahia K000004817
Carvalho, A.M. [1342], Bahia K000004818
Hage, J.L. [1734], Bahia K000004819
Wight, R. [1665], India K000245391 Unknown type material
Spruce, R. [5818], Ecuador K000201726
Spruce, R. [5818], Ecuador K000201727
Spruce, R. [5818], Ecuador K000201728
Hinton, G.B. [12066], Michoacán K000476299
Philcox, D. [3172], Brazil K001075832
Sellow [s.n.], Brazil K001075825
s.coll. [Cat. no. 8654], India K001131127
Bourgeau, M. [3140], Mexico K000643125
Hinton, G.B. [1160], México State K000476301
s.coll. [Cat. no. 8654] K001131125
Laegaard [21791], Thailand K000688373
Dombrowski, L.T.D. [2419], Brazil K001075828
Angus [168], Brazil K001075826
Irwin, H.S. [5653], Brazil K001075830
Hitchcock, A.S. [13801], Hawaii K001056124 Unknown type material
Glaziou, A.F.M. [6954], Brazil K001075827
Lechler, P. [1545], Peru K001044597
s.coll. [Cat. no. 8654] K001131124
Spruce, R. [s.n.], Brazil K001075831
Hinton, G.B. [4800], México State K000476298
Hinton, G.B. [12066], Michoacán K000476300
s.coll. [Cat. no. 8654] K001131126
Harger, E.B. [426], Connecticut K001056123 isolectotype
Mueller, F. [473], Brazil K001075829

First published in Sp. Pl.: 1050 (1753)

Accepted by

  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Aedo, C., Tellería, M.T. & Velayos, M. (eds.) (1999). Bases Documentales para la Flora de Guinea Ecuatorial; Plantas vascularis y hongos: 1-414. CSIC, real jardín Botánico, Madrid.
  • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
  • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
  • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
  • Bosser, J. & Renvoize, S.A. (2018). Flore des Mascareignes 203: 1-276. IRD Éditions, MSIRI, RBG-Kew, Paris.
  • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
  • Boulos, L. (2005). Flora of Egypt 4: 1-617. Al Hadara Publishing, Cairo.
  • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
  • Clayton, W.D. & Renvoize, S.A. (1982). Gramineae Flora of Tropical East Africa 3: 451-898.
  • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
  • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Cope, T.A., Knees, S.G. & Miller, A.G. (2007). Flora of the Arabian peninsula and Socotra 5(1): 1-387. Edinburgh University Press.
  • Danin, A. (2004). Distribution Atlas of Plants in the Flora Palaestina area: 1-517. The Israel Academy of Sciences and Humanities, Jerusalem.
  • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
  • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
  • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
  • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
  • Fosberg, F.R. & Renvoize, S.A. (1980). The Flora of Aldabra and neighbouring islands: 1-358. Crown, London.
  • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
  • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
  • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
  • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
  • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
  • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
  • Levin, G.A. & Moran, R. (1989). The vascular flora of isla Socorro, Mexico: 1-71. Dept. of Botany, San Diego Natural History Museum, San Diego.
  • Middleton, D.J. & al. (eds.) (2019). Flora of Singapore 7: 1-525. Singapore Botanic Gardens.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.
  • Patzelt, A., Harrison, T., Knees, S.G. & Hartley, L.A. (2014). Studies in the flora of Arabia: XXXI. New records from the Sultanate of Oman Edinburgh Journal of Botany 71: 161-180.
  • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
  • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
  • Robertson, S.A. (1989). Flowering Plants of Seychelles: 1-327. Royal Botanic Gardens, Kew.
  • Sosef, M.S.M. & al. (2019). Novitates Gabonenses 91: additons to the grass flora of Gabon Plant Ecology and Evolution 152: 99-105.
  • St. John, H. (1988). Census of the Flora of the Gambier islands, Polynesia Pacific Plant Studies 43: 1-34.
  • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
  • Veldkamp, J.F. (2014). A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes Blumea 59: 59-75.
  • Verloove, F. (2013). New xenophytes from Gran Canaria (Canary islands, Spain), with emphasis on naturalised and (potentially) invasive species Collectanea Botanica 32: 59-82.
  • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
  • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
  • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
  • Zuloaga, F.O., Morrone, O. & Pensiero, J.F. (2014). Gramineae VI Flora del Paraguay 45: 1-392. Editions des Conservatoire et Jardin botaniques de la ville de Genève.
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.

Literature

Catálogo de Plantas y Líquenes de Colombia

  • ColPlantA (2021). "ColPlantA. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.colplanta.org/"

Flora of West Tropical Africa

  • F.T.A. 9: 1082.
  • Sp. Pl. 1050 (1753)

Kew Backbone Distributions

  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Aedo, C., Tellería, M.T. & Velayos, M. (eds.) (1999). Bases Documentales para la Flora de Guinea Ecuatorial; Plantas vascularis y hongos: 1-414. CSIC, real jardín Botánico, Madrid.
  • Ahmed, Z.U. (ed.) (2008). Encyclopedia of Flora and Fauna of Bangladesh 12: 1-505. Asiatic Society of Bangladesh.
  • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
  • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
  • Bosser, J. (1969). Gramiées des pasturages et des cultures a Madagascar: 1-440. ORSTOM, Paris.
  • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286.
  • Britton, N. (1918). Flora of Bermuda: 1-585. Charles Scribner's Sons, New York.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
  • Clayton, W.D. & Renvoize, S.A. (1982). Gramineae Flora of Tropical East Africa 3: 451-898.
  • Clayton, W.D. & Snow, N. (2010). A key to Pacific Grasses: 1-107. Kew Publishing, Royal Botanic Gardens, Kew.
  • Cope, T.A., Knees, S.G. & Miller, A.G. (2007). Flora of the Arabian peninsula and Socotra 5(1): 1-387. Edinburgh University Press.
  • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
  • Danihelka, J. Chrtek, J. & Kaplan, Z. (2012). Checklist of vascular plants of the Czech Republic Preslia. Casopsi Ceské Botanické Spolecnosti 84: 647-811.
  • Danin, A. (2004). Distribution Atlas of Plants in the Flora Palaestina area: 1-517. The Israel Academy of Sciences and Humanities, Jerusalem.
  • Dassanayake (ed.) (1994). A Revised Handbook to the Flora of Ceylon 8: 1-458. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
  • Dobignard, D. & Chatelain, C. (2010). Index synonymique de la flore d'Afrique du nord 1: 1-455. Éditions des conservatoire et jardin botaniques, Genève.
  • Dávila, P., Mejia-Saulés, M.T., Gómez-Sánchez, N., Valdés-Reyna, J., Ortíz, J.J., Morín, C., Castrejón, J. & Ocampo, A. (2006). Catálogo de las Gramíneas de México: 1-671. CONABIO, México city.
  • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
  • Forzza, R.C. & al. (2013). Lista de Espécies da Flora do Brasil http://floradobrasil.jbrj.gov.br/2010/.
  • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
  • Giraldo-Cañas, D. (2011). Catálogo de la familia Poaceae en Colombia Darwiniana 49: 139-247.
  • Greuter, W. & Raus, T. (eds.) (2012). Med-Checklist Notulae, 31 Willdenowia 42: 287-295.
  • Henty, E.E. (1969). A manual of the grasses of New Guinea: 1-215. Division of Botany, Dpt. of Forests, LAE, New Guinea.
  • Hepper, F.N. (ed.) (1972). Flora of West Tropical Africa, ed. 2, 3(2): 277-574.
  • Hubbard, C.E. & Vaughan, R.E. (1940). Grasses of Mauritius and Rodriguez: 1-128. Crown Agents for the Colonies, London.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
  • Koyama, T. (1987). Grasses of Japan and its neighboring regions: an identification manual: 1-570. Kodansha, Tokyo, Japan.
  • Lebrun, J.P. (1973). Énumération des plantes vasculaires du Sénégal: 1-209. Maisons Alfort: Institut d'élevage et de médecine vétérinaire des pays tropicaux.
  • Levin, G.A. & Moran, R. (1989). The vascular flora of isla Socorro, Mexico: 1-71. Dept. of Botany, San Diego Natural History Museum, San Diego.
  • Masamune, G. (1942). Enumeratio phanerogamarum Bornearum: 1-739. Taihoku, Formosa.
  • Morat, P. & Veillon, J.-M. (1985). Contributions à la conaissance de la végétation et de la flore de Wallis et Futuna Bulletin du Muséum National d'Histoire Naturelle. Section B, Adansonia 7: 259-329.
  • Parham, B.E.V. (1971). The Vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll New Zealand Journal of Botany 9: 576-609.
  • Patzelt, A., Harrison, T., Knees, S.G. & Hartley, L.A. (2014). Studies in the flora of Arabia: XXXI. New records from the Sultanate of Oman Edinburgh Journal of Botany 71: 161-180.
  • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
  • Renvoize, S.A. (1998). Gramineas de Bolivia: 1-644. The Royal Botanic Gardens, Kew.
  • Robertson, S.A. (1989). Flowering Plants of Seychelles: 1-327. Royal Botanic Gardens, Kew.
  • Sheppard, C.R.C. & Seaward, M.R.D. (eds.) (1999). Ecology of the Chagos archipelago: 1-350. Westbury Academic & Scientific Publishing, Otley.
  • Sosef, M.S.M. & al. (2019). Novitates Gabonenses 91: additons to the grass flora of Gabon Plant Ecology and Evolution 152: 99-105.
  • Veldkamp, J.F. (2014). A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes Blumea 59: 59-75.
  • Verloove, F. (2013). New xenophytes from Gran Canaria (Canary islands, Spain), with emphasis on naturalised and (potentially) invasive species Collectanea Botanica 32: 59-82.
  • Walker, E.H. (1976). Flora of Okinawa and the southern Ryukyu islands: 1-1159. Smithsonian Institution Press, Washington D.C., U.S.A.
  • Woodroffe, C.D. (1985). Vegetation and flora of Nui Atoll, Tuvalu Atoll Research Bulletin 283: 1-18.
  • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348.
  • de Cordemoy, E.J. (1895). Flore de l'ile de la Réunion: 1-574. Librairie des sciences naturelles Paul Klincksieck, Paris.

Flora of Tropical East Africa

  • Clayton in Flora of West Tropical Africa, ed. 2, 3: 464 (1972).
  • L., Sp. Pl.: 1050 (1753).
  • Stapf & C. E. Hubbard in Flora of Tropical Africa 9: 1082 (1934).

Universidad Nacional de Colombia

  • Instituto de Investigación de Recursos Biológicos Alexander von Humboldt. (2014). Plantas alimenticias y medicinales nativas de Colombia. 2567 registros, aportados por: Castellanos, C. (Contacto del recurso), Valderrama, N. (Creador del recurso, Autor), B

Catálogo de Plantas y Líquenes de Colombia
http://creativecommons.org/licenses/by-nc-sa/3.0

Colombian resources for Plants made Accessible
http://creativecommons.org/licenses/by/3.0/

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of West Tropical Africa
Flora of West Tropical Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

GrassBase - The Online World Grass Flora
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Science Photographs
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Universidad Nacional de Colombia
ColPlantA database
http://creativecommons.org/licenses/by-nc-sa/3.0