1. Family: Lamiaceae Martinov
    1. Ajuga L.

      1. This genus is accepted, and its native range is Old World.

    [FTEA]

    Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Ajuga, C.W. Lukhoba. Flora of Tropical East Africa. 2009

    Habit
    Perennial, annual and biennial herbs (rarely subshrubs)
    Inflorescences
    Inflorescence of dense, verticillate cymes, sometimes reduced to solitary flowers or merging to form a terminal, spiciform thyrse
    Calyx
    Calyx actinomorphic or weakly zygomorphic (posterior lobe reduced), not accrescent, 5-lobed
    Corolla
    Corolla highly variable in colour, 2-lipped, marcescent, persisting as a sheath around fruit; posterior lip short, much shorter than anterior, 2-lobed, from which the sex organs emerge; anterior lip 3-lobed, median lobe very large, broadening distally and usually emarginate or bipartite
    Stamens
    Stamens 4, didynamous, usually slightly exserted; filaments fused to corolla for more than half their length, and covered with short hyaline hairs all over; anther thecae divergent to divaricate, confluent at dehiscence
    Disc
    Disk poorly to well developed, usually asymmetrical
    Ovary
    Ovary shallowly 4-lobed; style subterminal, ± equalling filaments, bifid with equal or unequal lobes
    Fruits
    Fruit usually a schizocarp of 4 nutlets (rarely drupaceous, 4-lobed); nutlets obovoid, reticulately foveolate, glabrous, with prominent lateral attachment-scar; endosperm present.
    Distribution
    Range: A genus of about 30 species, mostly tropical and subtropical in the Old World.
    [LKGF]

    Harley, R.M. et al. (2004). Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Springer-Verlag, Berlin, Heidelberg, New York.

    Habit
    Perennial, annual, and biennial herbs (rarely subshrubs)
    Inflorescences
    Inflorescence of dense, verticillastrate cymes, sometimes reduced to solitary flowers or merging to form a terminal, spiciform or subcapitate thyrse
    Calyx
    Calyx actinomorphic or weakly zygomorphic (posterior lobe reduced), not accrescent, 5-lobed
    Corolla
    Corolla highly variable in colour, 2-lipped or 1-lipped and then with only anterior lip present, marcescent, persisting as a sheath around fruit, posterior lip short or obsolescent, usually 2-lobed or retuse, anterior lip 3-lobed, median lobe very large,
    Stamens
    Stamens 4, didynamous, usually slightly exserted, thecae divergent to divaricate, confluent at dehiscence
    Pollen
    Pollen suprareticulate, columellae simple or branched
    Ovary
    Ovary shallowly 4-lobed
    Stigma
    Stigma lobes equal to unequal
    Disc
    Disc poorly to well developed, usually asymmetrical
    Fruits
    Fruit usually a schizocarp of 4 nutlets (rarely drupaceous, 4-lobed), nutlets oblate to obovoid, reticulately ridged to foveolate, usually glabrous, attachment-scar lateral, 0.55-0.85x as long as nutlet
    Endosperm
    Endosperm present.
    Note
    2n = 32 in most species (16, 24, 28, 30, 62, 64, ca. 86).  Forty to 50 species, paleotemperate, mostly Eurasian.
    Distribution
    Native to: Europe, Northern Africa, Macaronesia, West-Central Tropical Africa, Northeast Tropical Africa, East Tropical Africa, Southern Africa (doubtfully present in South Tropical Africa), Western Indian Ocean, Siberia, Russian Far East, Middle Asia, Ca
    [LKGF]
    Use
    Ajuga reptans L. is used as an ornamental ground cover.

    Images

    Distribution

    Native to:

    Afghanistan, Albania, Algeria, Amur, Assam, Austria, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bulgaria, Canary Is., Cape Provinces, Cape Verde, Central European Rus, China North-Central, China South-Central, China Southeast, Chita, Corse, Cyprus, Czechoslovakia, Denmark, East Aegean Is., East European Russia, East Himalaya, Egypt, Eritrea, Ethiopia, Finland, France, Free State, Germany, Great Britain, Greece, Hainan, Hungary, Iceland, India, Inner Mongolia, Iran, Iraq, Ireland, Italy, Japan, Kazakhstan, Kenya, Kirgizstan, Korea, Kriti, Krym, Kuril Is., KwaZulu-Natal, Laos, Lebanon-Syria, Lesotho, Libya, Madagascar, Madeira, Maluku, Manchuria, Morocco, Myanmar, Nansei-shoto, Nepal, Netherlands, New Guinea, New South Wales, North Caucasus, North European Russi, Northern Provinces, Northwest European R, Norway, Ogasawara-shoto, Pakistan, Palestine, Philippines, Poland, Portugal, Primorye, Qinghai, Queensland, Romania, Sardegna, Saudi Arabia, Sicilia, Sinai, South Australia, South European Russi, Spain, Sudan, Sulawesi, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tasmania, Thailand, Tibet, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Uganda, Ukraine, Victoria, Vietnam, West Himalaya, Western Sahara, Yemen, Yugoslavia, Zaïre

    Introduced into:

    Alabama, Arkansas, British Columbia, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Irkutsk, Kansas, Kentucky, Louisiana, Maine, Maryland, Masachusettes, Michigan, Minnesota, Mississippi, New Hampshire, New Jersey, New York, New Zealand North, New Zealand South, Newfoundland, North Carolina, Nova Scotia, Ohio, Ontario, Oregon, Pennsylvania, Québec, Rhode I., South Carolina, Tennessee, Texas, Utah, Venezuela, Vermont, Virginia, Washington, West Virginia, Wisconsin

    Ajuga L. appears in other Kew resources:

    First published in Sp. Pl.: 561 (1753)

    Accepted by

    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1995). World Checklist of Seed Plants 1(1, 2): 1-483, 1-529. MIM, Deurne.

    Literature

    Flora of Tropical East Africa
    • Kubitzki, Fam. Gen. Vasc. Pl. 7: 201 (2004)
    • Gen. Pl., ed. 5: 246 (1754)
    • Sp. Pl.: 561 (1753)
    Lamiaceae Key Genus Fact Sheets
    • Harley RM, Atkins S, Budantsev AL, Cantino PD, Conn BJ, Grayer R, Harley MM, de Kok RPJ, Krestovskaja T, Morales R, Paton AJ, Ryding O, and Upson T. 2004. Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Sp
    • Bulga Kuntze, Revis. Gen. Pl. 2: 512 (1891).
    • Rosenbachia Regel, Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 613 (1886).
    • Abiga St.-Lag., Ann. Soc. Bot. Lyon 7: 85 (1880).
    • Phleboanthe Tausch, Flora 11: 322 (1828).
    • Moscharia Forssk., Fl. Aegypt.-Arab.: 158 (1775).
    • Chamaepitys Hill, Brit. Herb.: 371 (1756).
    • Gen. Pl. ed. 5: 246 (1754).
    • Bugula Mill., Gard. Dict. Abr. ed. 4 (1754).
    • Ajuga L., Sp. Pl. 2: 561 (1753)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Lamiaceae Key Genus Fact Sheets
    Nina Davies, Gemma Bramley and Don Kirkup, Royal Botanic Gardens, Kew
    http://creativecommons.org/licenses/by/3.0