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This species is accepted, and its native range is Bioko to Somalia and S. Africa, Madagascar, Réunion, Himalaya to China (Yunnan), Taiwan.


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Bioko [Fernando Po], Clarence Peak, Mann 380 (K!, holo.)
Morphology General Habit
Terrestrial or lithophyte.
Vegetative Multiplication Rhizomes
Rhizome erect or short-creeping, 2–4 cm wide and deep; scales dark brown-black, to 10 × 2 mm, mixed with filiform scales of same length or smaller, leaving stubble when breaking off; in upper part of stipe fewer and smaller, linear with wider base, to 2–4 mm, frequently with glandular tip
Morphology Leaves
Fronds tufted, 6–9 per plant, erect and spreading.
Morphology Leaves Stipes
Stipe 18–60+ cm long, 2–5 mm in diameter at base, with adaxial groove and lateral grooves, with dark brown-black linear scales 2–10 × 2 mm, base widened, often gland-tipped.
Morphology Leaves Leaf lamina
Lamina thin, pale to mid-green, red when very young, ovate to deltoid, 34 × 34 cm, 2-pinnate to 3-pinnatisect.
Morphology Leaves Rachis
Rachis pale brown to stramineous, sparsely scaly, but with tuft at junction with pinnae.
Morphology Leaves Pinnae
Pinnae in 10–14 pairs, gradually merging into pinnatifid tip marked by decurrent lamina; pinnules in 8–12 pairs, opposite or sub-opposite, lower ones with lobed or crenate margin, basal acroscopic segment always more deeply cut or extended along costa; mid-vein in pinnule with ± 2 pairs of veinlets, not reaching margin; abaxial lamina glabrous, veins and veinlets with few relatively large multiseptate hairs (0.5)–1 mm long
Morphology Reproductive morphology Sori
Sori frequently constrained to top half, rarely present throughout pinnule, 1–1.4 mm in diameter; indusium 0.8–1 mm in diameter, with a plain, heavily undulating margin, deep notch to its centre aligned with the underlying veinlet, backward-pointing to the next larger vein or parallel to costa, stalk 0.3 mm long, with thick, knobbly radial walls
Morphology Reproductive morphology Spores
Spores monolete, oval, 0.05 mm long, 0.03 mm wide, with widely spaced, coarse tuberculae
Figure 12
Moist forest, often near water, or bamboo forest; 1850–2950 m
Widespread; least concern (LC)
Ching states that his new genus occupies an intermediate systematic position between Dryopteris, Peranema and Diacalpe. The genus agrees with Peranema D.Don and DiacalpeBlume in habit, leaf texture and colour, pattern of pinnation and particularly in the presence on costa and costules above of peculiar thick, stout, rufo-brown setae at the insertion of veins and veinlets, but differs above all in the dryopteroid indusium, the position of which is superior in relation to the sorus.
Flora districts: U2 K3, K4 T2 T6 T7 Range: From Bioko and Cameroon to Ethiopia and south to South Africa; Madagascar, Asia


None recorded for our area

Native to:

Cameroon, Cape Provinces, China South-Central, East Himalaya, Equatorial Guinea, Gulf of Guinea Is., Kenya, KwaZulu-Natal, Madagascar, Nepal, Northern Provinces, Rwanda, Réunion, Somalia, Sudan, Taiwan, Tanzania, Tibet, Uganda, Zimbabwe

Dryopteris squamiseta (Hook.) Kuntze appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Dec 5, 2007 Wallich, N. [372], Nepal Nothoperanema squamisetum K001080842
Mann, G. [380], Equatorial Guinea Nothoperanema squamisetum K000351166
Mann, G. [380], Equatorial Guinea Nothoperanema squamisetum K000351167
Buchanan [108], South Africa Nothoperanema squamisetum K000351168
Buchanan [108], South Africa Nothoperanema squamisetum K000351169 Unknown type material
Buchanan [20], South Africa Nothoperanema squamisetum K000351170

First published in Revis. Gen. Pl. 2: 813 (1891)

Accepted by

  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Kew Backbone Distributions

  • Fraser-Jenkins, C.R. & Kandel, D.R. (2019). Ferns and Fern-Allies of Nepal 2: 1-446. Government of Nepal, Ministry of Forests and Environment, Department of Plant Resources.

Flora of Tropical East Africa
Flora of Tropical East Africa

Herbarium Catalogue Specimens
'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
Digital Image © Board of Trustees, RBG Kew

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.