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This species is accepted, and its native range is Tropical Africa, Madagascar.


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Bioko [Fernando Po], on the peak, 2000 ft, 1860, Mann s.n. (K!, holo.)
Morphology General Habit
Vegetative Multiplication Rhizomes
Rhizome erect to short-decumbent, mostly unbranched, to 8 mm in diameter, with closely spaced stipe bases and brown to ferrugineous subulate scales to 22 × 3 mm, closely set with short teeth
Morphology Leaves
Fronds tufted, 4–7 per plant, arching, to 1 m long.
Morphology Leaves Stipes
Stipe greenish to stramineous, up to 48 cm long and up to 5 mm in diameter, proximally densely scaly, higher up moderately scaly, scales stramineous to ferrugineous, up to 15 × 4.5 mm, basally frequently with filiform outgrowths, often with scattered glandular cells, denticulate.
Morphology Leaves Leaf lamina
Lamina herbaceous, ovate, up to 53 cm long, to 2-pinnate-pinnatifid, proliferous, generally with one or more scaly buds adaxially along rachis near lamina apex.
Morphology Leaves Rachis
Rachis greenish to stramineous, narrowly winged towards apex, with ferrugineous to stramineous filiform to lanceolate scales up to 7 × 1.8 mm, denticulate, basally with filiform outgrowths, higher up closely set with short teeth.
Morphology Leaves Pinnae
Pinnae in up to 12 stalked pairs, basal pinna pair mostly longest, inaequilaterally ovate, narrowly ovate to oblong-acuminate towards lamina apex, basiscopically developed, up to 250 × 95 mm, with up to 3 stalked pinnule pairs; pinna-rachis narrowly winged for most of length, abaxially with sparse to moderate stramineous to ferrugineous filiform to narrowly oblong scales up to 5 × 1 mm.
Morphology Leaves Pinnules
Pinnules symmetric or inaequilaterally narrowly ovate to ovate, basiscopically decurrent, pinnatifid, often basiscopically developed, acroscopic pinnule on basal pinnae up to 50 × 17 mm, basiscopic pinnule on basal pinnae up to 56 × 23 mm, lobes broadly oblong-obtuse, shallowly lobed, dentate, glabrous adaxially, abaxially sparsely set with hairs and scales up to 2 mm long along veins
Morphology Reproductive morphology Sori
Sori inframedial on vein branches, discrete at maturity, circular, to 1.5 mm in diameter; sporangium stalk simple or haired; exindusiate
Morphology Reproductive morphology Spores
Spores with short and long ridges, ruminate, 40–58 × 26–32 mm.
Figure 13: 3
Terrestrial in moist forests and riparian forests; (1000–)1450–1850(–2250) m
Widespread; least concern (LC)
Vida, in Helv. Chim. Acta 56: 2130 (1973) report the species as tetraploid: 2n = ± 164.
Flora districts: U2 K4 K5 K7 T3 T5 T6 T7 Range: Sierra Leone, Liberia, Ivory Coast, Nigeria, Bioko, Cameroon, Malawi, Mozambique and Zimbabwe; Madagascar


Aspidiaceae, A. H. G. Alston. Flora of West Tropical Africa. Ferns and Fern-Allies Supplement. 1959

At 2, 000 to 4, 400 ft. alt.


None recorded for our area

Native to:

Burundi, Cameroon, Equatorial Guinea, Guinea, Gulf of Guinea Is., Ivory Coast, Kenya, Liberia, Madagascar, Malawi, Mozambique, Nigeria, Rwanda, Sierra Leone, Tanzania, Uganda, Zimbabwe

Dryopteris manniana (Hook.) C.Chr. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Oct 17, 2001 Chase, N.C. [5705], Zimbabwe K000994002 holotype
Oct 17, 2001 Adam, J.G. [21180], Liberia K000994003
Aug 1, 2001 Etuge, M. [4406], Cameroon K000109134
Aug 1, 2001 Ryan, P. [418], Cameroon K000109075
Aug 1, 2001 Balding, S. [60], Cameroon K000108987
May 1, 1997 Etuge, M. [1358], Cameroon K000108988
May 1, 1997 Etuge, M. [1356], Cameroon K000108954
Mann, G. [351], Equatorial Guinea K000675774 Unknown type material
Mann, G. [s.n.], Equatorial Guinea K000675775
Kamau [588], Kenya K000696577

First published in Index Filic.: 27 (1905)

Accepted by

  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Flora of West Tropical Africa

  • Tard. in Mém. I.F.A.N. 28: 149, t. 29, fig. 4, 5 (1953).
  • Ind. Fil. 276 (1905)

Flora of Tropical East Africa

  • J.P. Roux, Conspect. southern Afr. Pterid.: 125 (2001).
  • U.K.W.F.: 36 (1994);
  • Burrows, S. Afr. Ferns: 303, t. 50/ 6, fig. 70/ 307, map (1990);
  • B.J.B.B. 55: 163 (1985);
  • Jacobsen, Ferns S. Afr.: 440, t. 331, map 161 (1983)
  • Schelpe & Diniz, Fl. Moçamb., Pterid.: 240 (1979)
  • Schelpe, F.Z., Pterid.: 223, t. 63 (1970)
  • Alston, Ferns W.T.A.: 70 (1959)
  • Mém. I.F.A.N. 28: 149, t. 29/ 4–5 (1950)
  • Ind. filic.: 276 (1905)

Flora of Tropical East Africa
Flora of Tropical East Africa

Flora of West Tropical Africa
Flora of West Tropical Africa

Herbarium Catalogue Specimens
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Kew Backbone Distributions
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Kew Names and Taxonomic Backbone
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© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.