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This genus is accepted, and its native range is Tropics & Subtropics.
A specimen from Kew's Herbarium


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Morphology General Habit
Medium sized to large ferns, terrestrial or epilithic
Vegetative Multiplication Rhizomes
Rhizome erect, even trunklike, with clathrate scales
Morphology Leaves Stipes
Stipes with adaxial groove and basal scales; blade oblong-lanceolate, pinnate-pinnatifid to tri-pinnate, catadromous, maybe anadromous at very base; penultimate divisions often oblique with unequal base, decurrent and wing-connected, margin entire, serrate, crenate to pinnatifid.
Morphology Leaves
Veins free, simple or forked, basal vein of a segment rarely springing from costa instead of costule. Fronds tufted.
Morphology General Indumentum
Indumentum of narrow paleae that proximally bear short marginal outgrowths and often unicellular glands, pluricellular and acicular hairs on axes and adaxial lamina surfaces occasionally with ctenitoid hairs
Morphology Reproductive morphology Sori
Sori small circular, median, on enclosed veinlets, indusiate; sporangium short-stalked
About 150 species in the tropics and south temperate area; absent from Australia

Native to:

Angola, Argentina Northeast, Argentina Northwest, Assam, Bahamas, Bangladesh, Belize, Bermuda, Bismarck Archipelago, Bolivia, Borneo, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burundi, Cameroon, Central African Repu, China South-Central, China Southeast, Colombia, Comoros, Costa Rica, Cuba, Dominican Republic, East Himalaya, Ecuador, El Salvador, Ethiopia, Fiji, Florida, French Guiana, Galápagos, Ghana, Guatemala, Guinea, Gulf of Guinea Is., Guyana, Hainan, Haiti, Hawaii, Honduras, Ivory Coast, Jamaica, Japan, Jawa, Kazan-retto, Kenya, Korea, Leeward Is., Lesser Sunda Is., Louisiana, Madagascar, Malawi, Malaya, Maluku, Marianas, Mauritius, Mexican Pacific Is., Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, Nansei-shoto, New Guinea, Nicaragua, Norfolk Is., Ogasawara-shoto, Panamá, Paraguay, Peru, Philippines, Pitcairn Is., Puerto Rico, Rodrigues, Réunion, Samoa, Society Is., Solomon Is., Sri Lanka, Sudan, Sulawesi, Sumatera, Suriname, Taiwan, Tanzania, Thailand, Tibet, Togo, Trinidad-Tobago, Tubuai Is., Uganda, Uruguay, Venezuela, Venezuelan Antilles, Vietnam, Windward Is., Zambia, Zaïre, Zimbabwe

Ctenitis (C.Chr.) C.Chr. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Beyrich [ s.n.], Brazil K000954441
La Croix, I.F. [4970], Congo K000501436

First published in Notul. Syst. (Paris) 7: 86 (1938)

Accepted by

  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Flora of Tropical East Africa

  • Dryopteris Adans. subgen Ctenitis C.Chr., Biol. Arb. Warming: 77 (1911)
  • Verdoorn, Man. Pteridol.: 543 (1938)

Flora of Tropical East Africa
Flora of Tropical East Africa

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.